The Ceteosaurus whose bones have been discovered in the upper beds of
the Great Oolite at Enslow Rocks, at the Kirtlington Railway Station,
north of Oxford, and some other places, was a species of Crocodile
nearly resembling the modern Gavial or Crocodile of the Ganges. This
huge whale-like reptile has been described by Professor John Phillips as
unmatched in size and strength by any of the largest inhabitants of the
Mesozoic land or sea--perhaps the largest animal that ever walked upon
the earth. A full-grown Ceteosaurus must have been _at least_ fifty feet
long, ten feet high, and of a proportionate bulk. In its habits it was,
probably, a marsh-loving or river-side animal, dwelling amidst filicene,
cycadaceous, and coniferous shrubs and trees full of insects and small
mammalia. The one small and imperfect tooth which has been found
resembles that of Iguanodon more than of any other reptile; and it seems
probable that the Ceteosaurus was nourished by vegetable food, which
abounded in the vicinity of its haunts, and was not obliged to contend
with the Megalosaurus for a scanty supply of more stimulating diet.[69]
[69] For a full account of the Ceteosaurus, see “The Geology of the
Thames Valley,” by Prof. John Phillips, F.R.S. 1871.
Another reptile allied to the Pterodactyle lived in this epoch--the
_Ramphorynchus_, distinguished from the Pterodactyle by a long tail. The
imprints which this curious animal has left upon the sandstone of the
period are impressions of its feet and the linear furrow made by its
tail. Like the Pterodactyle, the Ramphorynchus, which was about the size
of a crow, could not precisely fly, but, aided by the wing (a sort of
natural parachute formed by the membrane connecting the fingers with the
body), it could throw itself from a height upon its prey. Fig. 119
represents a restoration of this animal. The footprints in the soil are
in imitation of those which accompany the remains of the Ramphorynchus
in the Oolitic rocks, and they show the imprints of the anterior and
posterior feet and also the marks made by the tail.
This tail was very long, far surpassing in length the rest of the
vertebral column, and consisting of more than thirty vertebræ--which
were at first short, but rapidly elongate, retain their length for a
considerable distance, and then gradually diminish in size.
[Illustration: XVIII.--Ideal landscape of the Middle Oolitic Period.]
Another genus of Reptiles appears in the Middle Oolite, of which we have
had a glimpse in the Lias and Great Oolite of the preceding section.
This is the _Teleosaurus_, which the recent investigations of M. E.
Deslongchamps allow of re-construction. The Teleosaurus enables us to
form a pretty exact idea of these Crocodiles of the ancient seas--these
cuirassed Reptiles, which the German geologist Cotta describes as “the
great barons of the kingdom of Neptune, armed to the teeth, and clothed
in an impenetrable panoply; the true filibusters of the primitive seas.”
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