The country in which the Megatherium has been found is described by Mr.
Darwin as belonging to the great Pampean formation, which consists
partly of a reddish clay and in part of a highly calcareous marly rock.
Near the coast there are some plains formed from the wreck of the upper
plain, and from mud, gravel, and sand thrown up by the sea during the
slow elevation of the land, as shown by the raised beds of recent
shells. At Punta Alta there is a highly-interesting section of one of
the later-formed little plains, in which many remains of these gigantic
land-animals have been found. These were, says Mr. Darwin:--“First,
parts of three heads and other bones of the Megatherium, the huge
dimensions of which are expressed by its name. Secondly, the
_Megalonyx_, a great allied animal. Thirdly, the _Scelidotherium_, also
an allied animal, of which I obtained a nearly perfect skeleton: it must
have been as large as a rhinoceros; in the structure of its head it
comes, according to Professor Owen, nearest to the Cape Anteater, but in
some other respects it approaches to the Armadilloes. Fourthly, the
_Mylodon Darwinii_, a closely related genus, of little inferior size.
Fifthly, another gigantic edental quadruped. Sixthly, a large animal
with an osseous coat, in compartments, very like that of an armadillo.
Seventhly, an extinct kind of horse. Eighthly, a tooth of a
pachydermatous animal, probably the same with the Macrauchenia, a huge
beast with a long neck like a camel. Lastly, the Toxodon, perhaps one of
the strangest animals ever discovered; in size it equalled an Elephant
or Megatherium, but the structure of its teeth, as Professor Owen
states, proves indisputably that it was intimately related to the
Gnawers, the order which, at the present day, includes most of the
smallest quadrupeds; in many details it is allied to the pachydermata;
judging from the position of its eyes, ears, and nostrils, it was
probably aquatic, like the Dugong and Manatee, to which it is allied.
How wonderfully are the different orders--at the present time so well
separated--blended together in different points in the structure of the
Toxodon!”[101]
[101] “Journal of Researches,” &c., by Charles Darwin, p. 81.
[Illustration: Fig. 190.--Lower jaw of the Mylodon.]
The remains on which our knowledge of the _Scelidotherium_ is founded
include the cranium, which is nearly entire, with the teeth and part of
the os hyoides, seven cervical, eight dorsal, and five sacral vertebræ,
both the scapulæ, and some other bones. The remains of the cranium
indicate that its general form was an elongated slender compressed cone,
beginning behind by a flattened vertical base, expanding slightly to the
cheek-bone, and thence contracting to the anterior extremity. All these
parts were discovered in their natural relative positions, indicating,
as Mr. Darwin observes, that the gravelly formation in which they were
discovered had not been disturbed since its deposition.
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