Fishes seem to have been the master existences of two great geological
systems, mayhap of three, ere the age of reptiles began. Now, fishes
differ very much among themselves, some ranking nearly as low as worms,
some nearly as high as reptiles; and we find in the Old Red Sandstone
series of links which are wanting in the present creation, and the
absence of which occasions a wide gap between the two grand divisions of
fishes, the bony and the cartilaginous.
Of all the organisms of the system one of the most extraordinary is the
pterichthys, or winged fish, which the writer had the pleasure of
introducing to the acquaintance of geologists. Had Lamarck been the
discoverer he would unquestionably have held that he had caught a fish
almost in the act of wishing itself into a bird. There are wings which
want only feathers, a body which seems to have been as well adapted for
passing through the air as through water, and a tail with which to
steer.
My first idea regarding it was that I had discovered a connecting
link-between the tortoise and the fish. I submitted some of my specimens
to Mr. Murchison, and they furnished him with additional data by which
to construct the calculations he was then making respecting fossils, and
they added a new and very singular link to the chain of existence in its
relation to human knowledge. Agassiz confirmed the conclusions of
Murchison in almost every particular, deciding at once that the creature
must have been a fish.
Next to the pterichthys of the Lower Old Red Sandstone I shall place its
contemporary the coccosteus of Agassiz--a fish which in some respects
must have resembled it. Both were covered with an armour of thickly
tubercled bony plates, and both furnished with a vertebrated tail. The
coccosteus seems to have been most abundant. Another of the families of
the ichthyolites of the Old Red Sandstone--the cephalaspis--seems
almost to constitute a connecting link between fishes and crustaceans.
In the present creation fishes are either osseous or cartilaginous, that
is, with bony skeletons, or with a framework of elastic,
semi-transparent animal matter, like the shark; and the ichthyolites of
the Old Red Sandstone unite these characteristics, resembling in some
respects the osseous and in others the cartilaginous tribes. Agassiz at
once confirmed my suspicion that the ichthyolites of the Old Red
Sandstone were intermediate. Though it required skill to determine the
place of the pterichthys and coccosteus there could be no mistaking the
osteolepis--it must have been a fish, and a handsome one, too. But while
its head resembled the heads of the bony fishes, its tail differed in no
respects from the tails of the cartilaginous ones. And so through the
discovery of extinct species the gaps between existing species have been
bridged.
_III.--Place-Fixing in the Dim Past_
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