The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
As the cervical vertebræ of the Megatherium have their processes
comparatively weaker than in the Scelidotherium, and the anterior dorsal
spines are relatively shorter, it may be concluded, that whatever were
the extent or nature of the fossorial labours of the enormous claws with
which it was provided, the head did not co-operate with the digging
implements in their especial task in the same degree as in the
Scelidothere and Orycterope. At the same time there is no modification
of the cervical region of the spine of the Megathere corresponding with
those which we have seen to be subservient to the arboreal habits of the
sloth, a remark which will not be deemed superfluous by those who have
perused the acute observations and arguments adduced by M. Lund in
favour of the scansorial character of the extremities of the Megatherium
and Megalonyx.
The fragments of the dorsal vertebræ and ribs of the Scelidotherium,
which are figured in Plate XX., offer no modifications which need detain
our attention; they closely conform, excepting in the greater relative
height of the anterior dorsal spines, already noticed, with the
Megatherioid type. The sacrum manifests in its vast expanse, the great
development of the posterior transverse processes to join the ischium,
the capacious medullary cavity, and wide nervous foramina, a like
conformity with the Megatherium, and a corresponding harmony with the
disproportionate bulk of the hind legs.
BONES OF THE EXTREMITIES.
The Scapula in its double spine, the osseous arch formed by the
confluence of the acromion with the coracoid process, and the
substitution of a distinct foramen for the suprascapular notch, agrees
with that of the Megatherium: but the span of the acromial arch is
relatively wider, and the surface for the articulation of the clavicle
is better marked. This articular surface, which is distinctly shewn upon
the acromion of both the scapulæ in Pl. XX. is the more interesting, as
being the only evidence of the clavicle of the Scelidothere which we at
present possess; but it is enough to prove that this quadruped enjoyed
all the advantages in the actions of the fore extremity, which arise out
of the additional fixation of the shoulder-joint afforded by the
clavicle—a bone which the extinct Megatherioids are the largest of the
mammiferous class to possess in a completely developed state. The form,
position, and aspect of the glenoid cavity for the humerus closely
correspond with the condition of the same part in the Megatherium. The
limits of the acromial and coronoid portions of the arch were still
defineable in the present skeleton, which indicates the nonage of the
individual in the unanchylosed condition of most of the epiphysiaes.
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