The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
Of the bones of the face there remain only portions of the malar,
lachrymal, palatine, and maxillaries. The chief peculiarities of the
malar bone have been already noticed: the breadth of the base of the
descending masseteric processes is two inches two lines; its termination
is broken off: the length of the ascending post-orbital process of the
malar cannot be determined from the same cause, but it is fortunate that
sufficient of this part of the cranium should have been preserved to
give this evidence of the affinities of the Scelidothere to the
Megathere. The malar bone is continued anteriorly, in a regular curve
forwards and upwards, to the lachrymal bone, and completes, with it, the
anterior boundary of the orbit: the size of the orbit is relatively
smaller than in the Orycterope, and still less than in the Ant-eaters:
here, however, we have merely an exemplification of the general law
which regulates the relative size of the eye to the body in the
mammalia. The malar bone does not extend so far forwards in front of the
orbit as in either the Orycterope or Armadillo; in the inclination,
however, with which the sides of the face converge forwards from the
orbits, the Scelidothere holds an intermediate place between the
Armadillos and Orycterope.
The lachrymal bone does not extend so far upon the face in the
Scelidothere as in the Orycterope; in which respect the Scelidothere
resembles more the Megathere. The foramen for the exit of the
infra-orbital nerve has the same situation near the orbit as in the
Megathere; its absolute distance from the anterior border of the orbit
is only half that in the Orycterope. The foramen is single in the
Scelidothere, as in the Orycterope; in the Megathere there are two or
three ant-orbital foramina. The vertical diameter of this foramen is
eight lines, the transverse diameter four lines. So much of the outer
surface of the superior maxillary bones as has been preserved, is smooth
and vertical. Each superior maxillary bone contains the sockets of five
teeth, occupying an antero-posterior extent of three inches seven lines,
(Pl. XXII. and XXIII. fig. 3). The posterior alveolus is situated just
behind the transverse line, extending across the anterior boundary of
the orbits; the remaining sockets of the molar series extend forwards
three inches in front of the orbits. In the Megatherium, the roots of
the five superior molars are all situated behind the anterior boundary
of the orbit: in the Orycteropus, on the contrary, the grinders are all
placed in advance of the orbit; so that the Scelidothere resembles that
species more than the Megathere in the relative location of the teeth.
The palatal interspace between the roots of the last molar tooth of each
series is eleven lines; the palate gradually though slightly widens, as
it advances forwards: the posterior margin of the palate is terminated
by an acute-angled notch. In the breadth of the bony palate the
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