Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
_Attractive grouping of Flowers._--To the same need of
conspicuousness the combination of so many individually small flowers
into heads and bunches is probably due, producing such broad masses
as those of the elder, the guelder-rose, and most of the Umbelliferæ,
or such elegant bunches as those of the lilac, laburnum, horse
chestnut, and wistaria. In other cases minute flowers are gathered
into dense heads, as with _Globularia_, _Jasione_, clover, and all
the Compositæ; and among the latter the outer flowers are often
developed into a ray, as in the sunflowers, the daisies, and the
asters, forming a star-like compound flower, which is itself often
produced in immense profusion.
_Why Alpine Flowers are so Beautiful._--The beauty of alpine flowers
is almost proverbial. It consists either in the increased size of the
individual flowers as compared with the whole plant, in increased
intensity of colour, or in the massing of small flowers into dense
cushions of bright colour; and it is only in the higher Alps, above
the limit of forests and upwards towards the perpetual snow-line
that these characteristics are fully exhibited. This effort at
conspicuousness under adverse circumstances may be traced to the
comparative scarcity of winged insects in the higher regions, and
to the necessity for attracting them from a distance. Amid the
vast slopes of debris and the huge masses of rock so prevalent in
higher mountain regions, patches of intense colour can alone make
themselves visible and serve to attract the wandering butterfly from
the valleys. Mr. Hermann Müller’s careful observations have shown,
that in the higher Alps bees and most other groups of winged insects
are almost wanting, while butterflies are tolerably abundant; and
he has discovered, that in a number of cases where a lowland flower
is adapted to be fertilized by bees, its alpine ally has had its
structure so modified as to be adapted for fertilization only by
butterflies.[23] But bees are always (in the temperate zone) far
more abundant than butterflies, and this will be another reason why
flowers specially adapted to be fertilized by the latter should be
rendered unusually conspicuous. We find, accordingly, the yellow
primrose of the plains replaced by pink and magenta-coloured alpine
species; the straggling wild pinks of the lowlands by the masses of
large flowers in such mountain species as _Dianthus alpinus_ and
_D. glacialis_; the saxifrages of the high Alps with bunches of
flowers a foot long as in _Saxifraga longifolia_ and _S. cotyledon_,
or forming spreading masses of flowers as in _S. oppositifolia_;
while the soapworts, silenes, and louseworts are equally superior
to the allied species of the plains.
[23] _Nature_, vol. xi. pp. 32, 110.
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