Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
Animal life is, on the whole, far more abundant and more varied
within the tropics than in any other part of the globe, and a great
number of peculiar groups are found there which never extend into
temperate regions. Endless eccentricities of form, and extreme
richness of colour are its most prominent features; and these
are manifested in the highest degree in those equatorial lands
where the vegetation acquires its greatest beauty and its fullest
development. The causes of these essentially tropical features are
not to be found in the comparatively simple influence of solar light
and heat, but rather in the uniformity and permanence with which
these and all other terrestrial conditions have acted; neither
varying prejudicially throughout the year, nor having undergone any
important change for countless past ages. While successive glacial
periods have devastated the temperate zones, and destroyed most of
the larger and more specialized forms which during more favourable
epochs had been developed, the equatorial lands must always have
remained thronged with life; and have been unintermittingly subject
to those complex influences of organism upon organism, which seem the
main agents in developing the greatest variety of forms and filling
up every vacant place in nature. A constant struggle against the
vicissitudes and recurring severities of climate must always have
restricted the range of effective animal variation in the temperate
and frigid zones, and have checked all such developments of form
and colour as were in the least degree injurious in themselves,
or which co-existed with any constitutional incapacity to resist
great changes of temperature or other unfavourable conditions. Such
disadvantages were not experienced in the equatorial zone. The
struggle for existence as against the forces of nature was there
always less severe,--food was there more abundant and more regularly
supplied,--shelter and concealment were at all times more easily
obtained; and almost the only physical changes experienced, being
dependent on cosmical or geological changes, were so slow, that
variation and natural selection were always able to keep the teeming
mass of organisms in nicely balanced harmony with the changing
physical conditions. The equatorial zone, in short, exhibits to us
the result of a comparatively continuous and unchecked development
of organic forms; while in the temperate regions, there have been
a series of periodical checks and extinctions of a more or less
disastrous nature, necessitating the commencement of the work of
development in certain lines over and over again. In the one,
evolution has had a fair chance; in the other it has had countless
difficulties thrown in its way. The equatorial regions are then,
as regards their past and present life history, a more ancient
world than that represented by the temperate zones, a world in
which the laws which have governed the progressive development of
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