Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
from the volcano before it was extinguished.
Not only are humming-birds found over the whole extent of America,
from Sitka to Tierra-del-Fuego, and from the level of the sea to the
snow-line on the Andes, but they inhabit many of the islands at a
great distance from the mainland. The West Indian islands possess
fifteen distinct species belonging to eight different genera, and
these are so unlike any found on the continent that five of these
genera are peculiar to the Antilles. Even the Bahamas, so close to
Florida, possess two peculiar species. The small group of islands
called Tres Marias, about sixty miles from the west coast of Mexico,
has a peculiar species. More remarkable are the two humming-birds of
Juan Fernandez, situated in the Pacific Ocean, four hundred miles
west of Valparaiso in Chili, one of these being peculiar; while
another species inhabits the little island Mas-afuera, ninety miles
further west. The Galapagos, though very little further from the
mainland and much more extensive, have no humming-birds; neither
have the Falkland islands, and the reason seems to be that both
these groups are deficient in forest, and in fact have hardly any
trees or large shrubs, while there is a great paucity of flowers
and of insect life.
_Humming-birds of Juan Fernandez as illustrating Variation and
Natural Selection._--The three species which inhabit Juan Fernandez
and Mas-afuera present certain peculiarities of great interest. They
form a distinct genus, Eustephanus, one species of which inhabits
Chili as well as the island of Juan Fernandez. This, which may be
termed the Chilian species, is greenish in both sexes, whereas
in the two species peculiar to the islands the males are red or
reddish-brown, and the females green. The two red males differ
very slightly from each other, but the three green females differ
considerably; and the curious point is, that the female in the
smaller and more distant island somewhat resembles the same sex
in Chili, while the female of the Juan Fernandez species is very
distinct, although the males of the two islands are so much alike. As
this forms a comparatively simple case of the action of the laws of
variation and natural selection, it will be instructive to see if we
can picture to ourselves the process by which the changes have been
brought about. We must first go back to an unknown but rather remote
period, just before any humming-birds had reached these islands. At
that time a species of this peculiar genus, Eustephanus, must have
inhabited Chili; but we must not be sure that it was identically the
same as that which is now found there, because we know that species
are always undergoing change to a greater or less degree. After
perhaps many failures, one or more pairs of the Chilian bird got
blown across to Juan Fernandez, and finding the country favourable,
with plenty of forests and a fair abundance of flowers and insects,
they rapidly increased and permanently established themselves on the
island.
Public-domain text, read in full here on John Shaqi.
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