The positions of the protoplasm which the vesicles fashion change
continually; only the grainless protoplasm of the _pseudopodia_ develops
no air. After long and fruitless efforts a manifest fatigue sets in; the
animal gives up the attempt for a time, and resumes it after an interval
of repose.
Engelmann, the discoverer of these phenomena, says (Pflüger’s Archiv für
Physologie, Bd. II.): “The changes in volume in all the vesicles of the
same animal are for the most part synchronous, effected in the same
manner, and of like size. There are, however, not a few exceptions; it
often happens that some of them increase or diminish in volume much
faster than others; sometimes one may increase while another diminishes;
all the changes, however, are throughout unquestionably intentional. The
object of the air-vesicles is to bring the animal into such a position
that it can take fast hold of something with its _pseudopodia_. When
this has been obtained, the air disappears without our being able to
discover any other reason for its disappearance than the fact that it is
no longer needed. . . . If we bear these circumstances in mind, we can
almost always tell whether an _arcella_ will develop air-vesicles or no;
and if it has already developed them, we can tell whether they will
increase or diminish . . . The _arcellæ_, in fact, in this power of
altering their specific gravity possess a mechanism for raising
themselves to the top of the water, or lowering themselves to the bottom
at will. They use this not only in the abnormal circumstances of their
being under microscopical observation, but at all times, as may be known
by our being always able to find some specimens with air-bladders at the
top of the water in which they live.”
If what has been already advanced has failed to convince the reader of
the hopelessness of attempting to explain instinct as a mode of conscious
deliberation, he must admit that the following considerations are
conclusive. It is most certain that deliberation and conscious
reflection can only take account of such data as are consciously
perceived; if, then, it can be shown that data absolutely indispensable
for the arrival at a just conclusion cannot by any possibility have been
known consciously, the result can no longer be held as having had its
source in conscious deliberation. It is admitted that the only way in
which consciousness can arrive at a knowledge of exterior facts is by way
of an impression made upon the senses. We must, therefore, prove that a
knowledge of the facts indispensable for arrival at a just conclusion
could not have been thus acquired. This may be done as follows: {111}
for, Firstly, the facts in question lie in the future, and the present
gives no ground for conjecturing the time and manner of their subsequent
development.
Public-domain text, read in full here on John Shaqi.
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