Vegetable Teratology: An Account of the Principal Deviations from the Usual Construction of PlantsMasters, Maxwell T. (Maxwell Tylden)
Science
Vegetable Teratology: An Account of the Principal Deviations from the Usual Construction of Plants
Masters, Maxwell T. (Maxwell Tylden)
Plants -- Abnormalities
Fig. 62, copied from Cramer, shows an instance of this kind in
_Delphinium elatum_, where not only is the thalamus prolonged, and the
carpels separated, but from the axils of some of the latter which have
assumed from the disunion of their margins somewhat of the appearance of
leaves, other flowering branches proceed--axillary prolification. If, on
the other hand, the carpels be few in number, and placed in a
verticillate manner, the axis then generally passes upwards without any
change in the form or position of the carpels being apparent, as in a
proliferous columbine, figured in the 'Linnean Transactions,' vol.
xxiii, tab. 34, fig. 5.
When a flower with the ovary naturally inferior or adherent to the calyx
becomes prolified, a change in the relative position of the calyx and
ovary almost necessarily takes place, the latter becoming superior or
detached from the calyx; this has been already alluded to in
_Umbelliferae_. In a species of _Campanula_ examined by me, the calyx was
free, the corolla double, the stamens with petaloid filaments, and in
the place of the pistil there was a bud consisting of several series of
green bracts, arranged in threes, and enclosing quite in the centre
three carpellary leaves detached from one another and the other parts of
the flower, and open along their margins, where the ovules were placed.
In other similar instances in the same species of _Campanula_, the
styles were present, forming below an imperfect tube which surrounded
the adventitious bud; in another, contrary to what occurs usually in
such cases, the ovary was present in its usual position, but surmounted
by a bud of leafy scales, enclosed within the base of a tube formed by
the union of the styles. A similar relative change in the position of
the calyx and the ovary takes place when the _Compositae_ are affected
with central prolification, or even in that lesser degree of change
which merely consists in the separation and disunion of the parts of the
flower, but which in these flowers appear to be, as it were, the first
stage towards prolification. I owe to the kindness of Professor Oliver a
sketch of a species of _Rudbeckia_? showing this detachment of the calyx
from the ovary. In a monstrous _Fuchsia_ that I have had the opportunity
of recently examining, the calyx was similarly detached from the ovary
simultaneously with the extension of the axis. Here the petals were
increased in number and variously modified, the stamens also; while in
the centre and at the top of the flower, conjoined at the base with some
imperfect stamens, was a carpel open along its ovuliferous margins. Such
instances as these seem to be the first stages of a change which,
carried out more perfectly, would result in the formation of a new bud
on the extremity of the prolonged axis.
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