Venoms: Venomous Animals and Antivenomous Serum-therapeuticsCalmette, A. (Albert)
Science
Venoms: Venomous Animals and Antivenomous Serum-therapeutics
Calmette, A. (Albert)
Antitoxins; Poisonous snakes; Toxins; Venom
Each gland is surrounded by a thick capsule of fibrous tissue, two
prolongations of which, the one anterior, the other posterior, keep it
in its place beneath the _masseter_ muscle. A portion of the latter
is inserted in the capsule itself, in such a way that when the snake
closes its jaws to bite, the gland is forcibly compressed and the
contained liquid is squeezed into its excretory duct.
Between the muscle and the envelope of the gland there is a serous
pouch, which enables the one to slide over the other.
The excretory duct runs along the outer side of the upper jaw,
and opens by a slit at the base of the poison-fang, with which it
inosculates at right angles in a little muscular mass forming a
_sphincter_.
In the normal position of repose, the poison-fang is always concealed
by a gingival fold of mucous membrane, in the substance of which are
buried a few fibres of the tendon of the internal pterygoid muscle.
When the latter contracts, the tooth is almost completely exposed, and
the efferent duct of the gland then assumes an oblique position, which
allows of the direct discharge of the venom through the canal which
runs along the greater portion of the length of the tooth.
When the poison-fangs are folded back in their sheath, the poisonous
secretion can escape freely into the buccal cavity by the slit situated
at the base of the fangs.
At the moment when the animal is about to bite, when it throws back its
head and opens its jaws, directing its fangs forwards, the muscles that
come into action (_masseters_, _temporals_, and _pterygoids_) compress
the glands on each side, and cause the venom to be expelled in a sudden
jet, as if by a sort of ejaculatory process. In the case of certain
species the venom may be projected to a distance of more than a yard.
The quantity of venom secreted by the glands varies greatly, according
to the length of time which has elapsed since the animal took its last
meal, and in accordance with a number of other conditions not very easy
to determine.
The Common Viper of Europe yields scarcely 10 centigrammes of poison,
while an adult Indian Cobra may excrete more than 1 gramme.
* * * * *
Freshly collected venom is a syrupy liquid, citron-yellow or slightly
opalescent white in colour.
When dried rapidly _in vacuo_ or in a desiccator over calcium chloride,
it concretes in cracked translucent lamellæ like albumin or gum arabic,
and thus assumes a crystalloid aspect. In this condition it may be kept
indefinitely, if protected from light, air, and moisture. It dissolves
again in water just as readily as albumin or dried serums.
Public-domain text, read in full here on John Shaqi.
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