Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
A few years ago the writer described a very slender, lizard-like
reptile about two feet in length from the Permian of Texas under
the name _Araeoscelis_, so named because of its slender legs. The
structure of both the skull and the skeleton of this reptile is now
quite satisfactorily known, so well known indeed that the accompanying
restoration (Fig. 62) has little that is conjectural about it, at
least so far as the form is concerned. The skull has a single, upper
temporal opening, quite like that of lizards, but the quadrate is
not loose below. And this is really what we should expect in the
ancestral lizards; and everything else of the skeleton, except perhaps
one character, is what would be expected. That one character is the
elongation of the cervical vertebrae, which are about twice the length
of the dorsal vertebrae following them. The cervical ribs are very
slender bones, articulating by a single head with the centrum only. In
these and other characters, so far as they are known, _Araeoscelis_
seems to agree with _Protorosaurus_, and both have very hollow bones.
[Illustration: FIG. 63.—Skeleton of _Pleurosaurus_. (After Lortet)]
PLEUROSAURUS
We may for the present be justified in maintaining the order
Protorosauria for those reptiles having a single, typically upper
temporal opening on each side, with a fixed quadrate, not including
the ichthyosaurs. It is not improbable, however, that when more is
known of the ancestors of the lizards, the whole group will find its
most natural place among the Squamata. This definition will include
a peculiar aquatic reptile that has been known for many years, but
which has been wrongly classed in the same family as _Sphenodon_, on
the purely gratuitous assumption that it has two temporal openings
on each side; we now know that it has but one. This reptile, known
scientifically as _Pleurosaurus_, was described originally by H. von
Meyer in 1843, but we are indebted to M. Lortet for a more precise
knowledge of the animal, and for the figure (Fig. 63) which is here
given of the skeleton. Not a few excellent skeletons are preserved in
the museums at Lyons and Munich. The specimen here figured, as actually
preserved, measures about three feet in length; a part of the tail is
missing, which is known from other specimens to have been remarkably
long.
The figures show clearly some of the remarkable aquatic adaptations of
the animal, especially the short neck, the very long and narrow body,
and the extraordinarily long and flattened tail. The head is elongate
triangular in shape, resembling very much that of the mosasaurs; and
the external nostrils are likewise situated remotely from the end
of the snout, as in the mosasaurs. The extremity of the snout has a
beak-like projection. The teeth are much longer, more pointed, and more
recurved than is the case with most land reptiles, indicating their use
for the capture and retention of slippery, quick-moving prey.
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