Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
_Aigialosaurs._—Within recent years a number of other lizards have
been made known from the Lower Cretaceous rocks of Dalmatia which
present most remarkable intermediate characters between the monitors,
dolichosaurs, and the mosasaurs, the famous sea-lizards of Upper
Cretaceous age. Some of these lizards had twelve or thirteen vertebrae
in the neck, while others had but seven—an unusually short neck
characteristic of the mosasaurs. These latter kinds, belonging to two
or three genera, are included in a distinct group. They were long and
slender, the head long and pointed. The teeth, conical and sharp, were
attached in shallow pits, quite as in the mosasaurs. The lower jaws
had a hinge just back of the teeth, as in the mosasaurs, of which the
only trace in modern lizards is found among the monitors. Still more
remarkable, though perhaps not so easily appreciated, is the shape of
the quadrate bone, with a broad flaring rim for the ear cavity, quite
unlike that of land lizards, but quite like that of the mosasaurs. In
fact, the very peculiar skull is almost identical with that of the true
sea-lizards. The body and tail also resemble those of the mosasaurs
more than those of the monitors, but there is a firm attachment of the
pelvis to the backbone, and the legs are long and lizard-like, though
not as long as those of land lizards. The feet were webbed in life, and
the toes have no claws, conclusively demonstrating their water habits.
The vertebrae indeed have the same peculiar articulations, called
zygosphenes, as in most of the mosasaurs. The largest aigialosaurs were
about six feet in length, that is, of about the size of the smallest
known mosasaurs.
[Illustration: FIG. 68.—_Clidastes_, an American mosasaur. Life
restoration.]
We have then in the aigialosaurs nearly every known intermediate
character that we could wish for in a connecting link between the
mosasaurs and the monitors, lizards that were equally at home on land
or in the water, and there can be scarcely a doubt that they were
either the direct ancestors or closely akin to the direct ancestors of
the strictly marine mosasaurs; and scarcely a doubt that they were the
descendants of the actual forbears of the modern monitors, which, as we
have seen, have acquired partial aquatic habits in escaping from their
enemies. The dolichosaurs we can now understand were a side branch
from these semiaquatic aigialosaurs which, specializing in another
direction, quickly came to grief, perhaps in competition with their
more agile and skilful short-necked kin.
Taking all these facts into consideration it seems best to unite
the monitors, dolichosaurs, and aigialosaurs into one group of the
Lacertilia, the Platynota, intermediate in place between the true land
lizards and the truly aquatic mosasaurs.
MOSASAURS
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