Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
These semiaquatic reptiles never grew very large—not more than four
or five feet in length; nor did they ever succeed in becoming fully
at home in the water, certainly no more so than our modern alligators
and crocodiles. They remained to the end of their comparatively brief
existence essentially land animals, probably seeking their food in the
water but fleeing to the land for protection and for the breeding of
their young. Their chief water adaptations are seen in the elongate
face and flattened swimming tail. Their legs remained essentially
terrestrial, and could have been of but little use in the water for
propulsion; the feet even were doubtfully webbed, or if so, not more
than are those of the alligator. Singularly, like the proganosaurs,
their ribs were heavy and stout, also suggesting bottom-crawling
habits, like those of the living Galapagos lizards. The skull was
lightly built, and the face was long and slender, like that of the
gavials and proganosaurs; but, like those of the former and unlike
those of the latter, the nostrils were situated at the extreme tip.
The hind legs were firmly attached to the body by the sacrum; and no
sclerotic bones of the eyes have been discovered. The neck was neither
unusually long nor unusually short. The body was probably covered with
horny scales.
[Illustration: FIG. 88.—Restoration of _Champsosaurus_.]
[Illustration: FIG. 89.—_Champsosaurus_; skull from above. (After
Brown.)]
To the student of paleontology these animals are of interest because of
the retention of several primitive traits which had long disappeared in
other known reptiles. While the vertebrae had ceased to be perforated
by the notochord, as in the early reptiles, they were still shallowly
biconcave. The first bone of the neck, the atlas, had changed but
little from that of their very ancient forbears of Permian times, and
the bones of the palate still retained numerous teeth scattered over
it, like those of the same Paleozoic ancestors. Most primitive and
old fashioned of all was the pelvis, which was so unlike that of all
known contemporary or later reptiles that, were a paleontologist to
see it without knowing whence it came, he would be almost sure to say
that it belonged to a Paleozoic, or at least a Triassic, reptile, and
not only to an early reptile but a very primitive one at that. This
peculiarity consists in the absence of any opening between the ischium
and pubis, which is characteristic of every living vertebrate with
legs. And these and other old-fashioned characters could not possibly
have been new developments; they must have existed in all the ancestors
of the Choristodera from Paleozoic to early Tertiary times, though not
a single other reptile is known to have possessed them, for the greater
part of this time. Perhaps when Asia and northwestern America have
been more thoroughly explored for vertebrate fossils, some of their
ancestors which perished on their great migration from the western to
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