Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
The collar-bones, or clavicles, are wanting in crocodiles; there is a
slender interclavicle; and the shoulder-blade and coracoid are well
developed (Fig. 102). The bones of the pelvis are loosely united with
each other as they are in most reptiles (Fig. 104). The pubes, the
anterior bones below, unlike those of all other reptiles, do not help
to form the acetabulum or socket for the articulation of the thigh
bone, nor do they articulate with each other. This single character
sharply distinguishes a crocodile from all other reptiles, living or
extinct, and is almost the only character that separates the order from
the dinosaurs, aside from the peculiar structure of the nasal passages
in the skull. On the under side of the body, connected with the front
end of the pubes, there are seven or eight pairs of abdominal ribs,
corresponding to the plastron of the turtles and similar to those of
the ichthyosaurs and plesiosaurs. The mosasaurs have no such ribs.
[Illustration: FIG. 102.]
[Illustration: FIG. 103.
FIG. 102.—Scapula (_sc_) and coracoid (_cor_) of gavial.
FIG. 103.—Front foot of crocodile: _u_, ulna; _r_, radius; _re_,
radiale; _ue_, ulnare; _p_, pisiform.]
Furthermore, the crocodiles differ from all other living reptiles in
having a four-chambered heart, like that of birds and mammals, that
is, a heart with two auricles and two ventricles. This more perfect
structure of the circulatory organs does not, however, insure at all
times a complete separation of the pure or arterial blood from the
impure or venous blood, since the blood may be more or less intermixed
outside of the heart by a connection between the venous and the
arterial systems. Whether these imperfectly developed organs, so
suggestive of a higher and more perfect mode of respiration, are the
vestiges of what were once among some reptiles functional structures,
or whether they are rudiments of a higher organization, developing
independently in these creatures, cannot be positively determined,
but it seems very probable that, far back in geological times, some
reptiles, especially the pterodactyls and dinosaurs, had their
respiratory and circulatory systems more like those of the birds and
mammals of today. Unfortunately, however, if such was the case, we may
never be able to prove it, although proof would not be impossible;
stranger things than fossil hearts have been found by paleontologists!
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