Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
We recognize in _Geosaurus_ an unusually slenderly
built crocodile, in appearance very different from all true
crocodiles. The smooth, rounded skull, with its greatly
elongated and slender snout, and the deep-lying, small eyes,
reminds one most of the ichthyosaurs. The skull merges into
the slender, elongated trunk without a visible neck, and
the body was provided neither above nor below with horny or
bony armor, but was, probably, as in the whales, covered
with a smooth, soft skin. The anterior extremities, attached
far forward, are developed as paddles, and served rather
as organs of equilibration than as a means of propulsion,
which was the function of the elongated hind legs and the
extraordinarily strong and powerful tail, which supported
at its end a large fin. The entire impression given of
the animal is that of an excellent swimmer, with all the
peculiar aquatic adaptations. In the skeleton, however, all
the characters of the original crocodiles are preserved.
Most remarkable are the laterally placed eyes, protected
by the stout sclerotic bones, and the overhanging bones of
the orbits. So, too, the large temporal openings of the
skull, doubtless due to the absence of the bony plates in
the integument, give to the animal a strangely abnormal
appearance for a crocodile.
We have observed that all the truly aquatic air-breathing animals,
save the plesiosaurs, have either lost the hind legs or else have them
greatly reduced in size, and the disproportionately large size of
these members in _Geosaurus_ seems inexplicable. But an explanation is
not, I think, hard to find. In the adaptation to water life the first
to become modified for the control of the body are the front legs.
The hind legs never have any really important use when the tail is a
powerful propeller. The hind legs of the geosaurs are still essentially
legs and not paddles, and they were doubtless used either occasionally
for propulsion on land, or perhaps for pushing the body about on
the bottom of shallow waters. And the presence of a well-developed
ventral armor of bony ribs possibly also indicates more or less of the
terrestrial crawling habit. As soon as the hind legs cease to be used
for crawling they take on only a feeble use for the equilibration of
the body, and speedily become small, until finally they disappear.
That the hind legs of these creatures were of some use in the water
is certain, because of the modifications in their structure, and
especially because of the loss of the claws; but that they were of
important use as propellers is hardly probable. We may therefore infer
that the thalattosuchians, while distinctively sea-reptiles, had not
entirely lost their land habits. Moreover, it is highly probable that
their egg-laying habits, which would hardly change with a greater
adaptation to water life, compelled the animals recurrently to visit
the shores.
Public-domain text, read in full here on John Shaqi.
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