Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
The Pleurodira, as the term indicates, are easily distinguished from
all other turtles by the way in which they withdraw the head within the
shell. Instead of withdrawing it by an =S=-shaped flexure of the neck
between the shoulder-blades, as do other turtles, these bend the neck
laterally in a horizontal plane, bringing the head within the margins
of the shell in front of one or the other foreleg; and the margins of
the shell are produced here in an eave-like fashion for the greater
protection of the head. In the structure of the shell, which is always
fully developed into a box, these turtles do not differ very much from
the Cryptodira, though there may be some extra bones in the plastron,
as also in the skull. The nasal bones are always, the lacrimals
sometimes, well developed; the latter never, the former rarely, found
in other groups. The lower jaws articulate a little differently,
and the external ear is always fully surrounded by bone. Very
characteristic is the bony union of the pelvis with the plastron below,
which never occurs in other turtles, unless it be the Amphichelydia.
The side-necked turtles are all of fresh-water habit, similar to that
of the fresh-water tortoises spoken of farther on. The neck is often
very long and snake-like, which accounts for one of the names given
to these turtles; because it is withdrawn into the shell sidewise, it
has more distinctively ball-and-socket joints between the vertebrae,
with distinct transverse processes for the attachment of the necessary
side-moving muscles. The feet in all are more or less webbed and armed
with strong claws.
The largest and perhaps the best known of all living side-necked
turtles is the giant Amazon turtle of South America, which sometimes
has a shell nearly three feet in length. Its feet are broadly webbed,
and the shell is rather flat in the adult; it is an excellent swimmer
in the waters of the Orinoco and Amazon. Six or seven species of
the genus to which it belongs are known, all of them South American
except one that lives in Madagascar and one fossil found in the Eocene
of India. This remarkable distribution is but one more of the many
instances known in zoölogy and paleontology that seem to prove an early
land connection between India and South America. Had the migration
between the two continents occurred by way of Asia and Bering straits,
as did that of hosts of mammals, one would certainly expect to find
some evidence of it in the North American Tertiary rocks, which, so
far, is lacking.
CRYPTODIRA
The chief families of the Cryptodira turtles are the Chelydridae,
or snappers; the Emydidae, or marsh tortoises; the Testudinidae, or
land tortoises; the Chelonidae, or sea-turtles; the Protostegidae, or
ancient sea-turtles; and the Dermochelydidae, or leather-backs. Other
doubtful or smaller groups, both living and extinct, may be omitted, or
incidentally mentioned.
SNAPPING TURTLES
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