Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
Certain it is that the common box tortoise, of like form and habits,
occurs not rarely on the arid plains, far from water.
The neck and legs became fully retractile within the shell; the digits
were shortened up, without a vestige of webbing membrane between them;
the phalanges were reduced in number to two in each toe, and nearly all
the toes have well formed claws. The feet are placed squarely upon the
ground, and the body is elevated in walking. They can swim, when by
accident they are thrown into the water, only as any terrestrial mammal
can.
About forty species of land tortoises are known throughout the world
at the present time, though North America, the probable original home
of the tribe, has but three, all small. The larger species are all now
denizens of islands, especially the Galapagos Islands, where the giant
tortoises have long been famous. And many of our living forms have
changed but little since Eocene times. In the Oligocene and Miocene
they inhabited western North America in enormous numbers. In the Bad
Lands of South Dakota one can often see the remains of a dozen or more
of these giant tortoises at one time, specimens varying from one to
three feet in length of shell. In river deposits, those of the late
Miocene or early Pliocene, the writer has seen areas of an acre or more
literally strewn with their remains, as though droves of them had been
overwhelmed and perished together. About fifty species of these land
tortoises are known from the American Tertiary, thirty-two of them
belonging to the modern genus _Testudo_, which comprises the giant
tortoises of the Galapagos. The largest known species of the group is
one of _Testudo_ from the Pliocene of India, which had a shell six feet
in length. Why the larger species became extinct in Pliocene times on
the mainland to survive only in the islands is not known; possibly
their carnivorous enemies became too cunning and too numerous.
SEA-TURTLES. CHELONIDAE
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