Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
In Fig. 38 on p. 85 are shown two paddles, the front and hind paddles
of a single individual of a very specialized plesiosaur from the
Upper Cretaceous of Kansas (_Trinacromerum_). The long arm and thigh
bones are followed by remarkably short and broad bones in place of
the elongated forearm and leg bones of the land reptiles. Not only
are these bones much broader than they are long, but there have been
developed additional bones back of them in the same row—new bones
which have no counterpart in any terrestrial reptiles. In the first of
the three figures is shown a hind paddle of one of the earliest known
plesiosaurs, _Thaumatosaurus_, from the lower part of the Jurassic of
Germany. It will be seen here that the tibia and fibula are much more
elongated than in _Trinacromerum_, and much more like the leg bones
of land reptiles. A still more primitive stage in the evolution of
the swimming paddle of the plesiosaurs will be seen in Fig. 48 on p.
99, the possibly ancestral, amphibious nothosaur. Here the tibia and
fibula, while relatively very much shorter than in any land reptile,
still have, together with all the other bones of the leg, a terrestrial
or amphibious type. In Fig. 39 is seen the front paddles of the
long-necked _Elasmosaurus_, which, though one of the latest of all
plesiosaurs in geological history, has the structure of its paddles
somewhat intermediate between that of the earlier _Plesiosaurus_ and
the later _Trinacromerum_.
[Illustration: FIG. 40.—Skull of _Elasmosaurus_ from the side: _pm_,
premaxilla; _m_, maxilla; _po_, postorbital; _j_, jugal.]
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