Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
The hind limbs were smaller,
often much smaller than the fore ones, and both were quite fin-like in
life, or rather flipper-like, though not at all fin-like in structure.
The skin was smooth and bare. In brief, to quote Fraas’s words:.
The general aspect of the ichthyosaurs was very
dolphin-like. The body was everywhere naked and probably
dark in color. The head was produced in front into a
long, slender snout, and was closely joined to the body
posteriorly without indications of a neck. The body itself
was cylindrical, expanded in front by the large thorax and
abdomen, but rapidly diminishing into the long, slender, and
strong tail. Close behind the head were the front paddles,
which in some species were broad and shovel-like, in others
elongated and pointed. The hind paddles were smaller than
the front ones, sometimes greatly reduced in size, their
function replaced by that of the very broad tail.
From the foregoing descriptions and the restoration shown in Fig. 51,
we see how very fish-like, or rather dolphin-like, these animals were
in the external form—so fish-like that the name _Ichthyosaurus_ is not
misleading, though Koenig gave it in the mistaken belief that they were
really allied to the fishes. When to these external features certain
other fish-like details of the skeleton are added, we do not wonder
that the early observers were so long in doubt about them. A more
careful examination of the skeleton will, however, disclose so many
truly reptilian characters that their external appearance and habits
lose all significance.
The vertebrae are deeply biconcave and fish-like, it is true, but a
consideration of the reasons therefor will convince us that any other
kind of vertebrae would be more remarkable. At the time when the
ichthyosaurs must have originated, at the time when the first known
ichthyosaurs appeared in geological history indeed, all reptiles had
biconcave vertebrae, and for the most part at least deeply biconcave
ones. The vertebrae remained fish-like throughout all their history,
perpetuating their type until most other reptiles had developed a
firmer one, because such vertebrae were best adapted for the quick,
pliant movements of the spinal column so necessary for the well-being
of the animals in the water. In the modern dolphins, animals in shape,
size, and habits most wonderfully allied to what these old reptiles
must have been, the small, flat-ended vertebrae are widely separated by
disks of flexible cartilage.
Not only were their vertebrae fish-like in form, but there are other
characters in the spinal column of a primitive or generalized nature.
As in all aquatic animals, the articulating processes between the
vertebrae are either weak or wanting in the posterior part of the
column. And they were not only small, but were situated, in many,
high up, very remarkably resembling the peculiar arrangement of the
articulations in the dolphins.
Public-domain text, read in full here on John Shaqi.
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