Water Reptiles of the Past and PresentWilliston, Samuel Wendell
Science
Water Reptiles of the Past and Present
Williston, Samuel Wendell
Aquatic reptiles; Aquatic reptiles, Fossil
Their vertebrae, as would be expected in such old reptiles, are quite
primitive in structure, that is, they are deeply concave in each end,
probably being perforated for the remains of the notochord. The pelvis
also is of the old-fashioned type, that is, without an opening or
vacuity between the bones below. The shoulder bones are old fashioned
too. The shoulder-blade, especially, shows a decided adaptation to
water life in its short, fan-like shape, very much like those of the
mosasaurs, ichthyosaurs, whales, etc. Just why swimming animals should
have short and broad shoulder-blades has not yet been explained, but
doubtless they afforded better attachment for those muscles used more
especially in swimming. The ribs are remarkably flat and heavy, and
were not very firmly attached to the vertebrae. Heavy ribs are unusual
among free swimming animals, but do occur in the modern sirenians,
which live on the bottoms of shallow bays, etc., feeding upon plants.
We may perhaps infer from this peculiar structure of the ribs that
the proganosaurs lived more on the bottoms of shallow waters, feeding
upon such fishes or invertebrates as they could capture, coming to the
surface to breathe from time to time. Possibly they sought the shores
for safety from their enemies, as do the Galapagos lizards, figured
on p. 142; and doubtless they laid and hatched their eggs on land.
A character which suggests that the proganosaurs lived only in the
shallow waters is the elongated neck, reminding one of those two other
groups of swimming reptiles, the dolichosaur lizards and the nothosaurs
of the Sauropterygia, the only known reptiles besides the plesiosaurs
having an abnormal number of neck bones. Still more suggestive of
shallow, fresh-water habits is the absence of eye bones, as in the
modern crocodiles.
The long snout, with the long and slender teeth, and the position
of the external nostrils far back near the eyes, together with the
flattened and long tail and the webbed feet, are sufficient proof of
expert swimming habits. The legs still functioned more or less for the
support and propulsion of the body on the land, and they probably were
only of slight service in the water. The alligator swims sinuously
with its front legs collapsed and extended by the side of the body;
its hind legs are used more as propellers, with the knee flexed and
the feet turned outward and expanded. The legs of the proganosaurs
doubtless were used in the same way, as shown in the restoration, which
has been modified from the original of Dr. McGregor in accordance with
this probable use of the legs.
Public-domain text, read in full here on John Shaqi.
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