[Illustration: Fig. 61. Mixed Hardwoods on Lower Levels. Spruce
and Balsam Dominate on Higher Elevations. Mt. McIntyre, Adirondack
Mountains, New York. _U. S. Forest Service._]
[Illustration: Fig. 62. Scrub Growth on Mountain Top. Mt. Webster, New
Hampshire. _U. S. Forest Service._]
LIGHT.
More than by any other factor, the growth of trees in a forest is
determined by the effect of light. All trees need light sooner or
later, but some trees have much more ability than others to grow
in the shade when young. Such trees, of which maple and spruce are
examples, are called _tolerant_, while others, for instance, larch,
which will endure only a comparatively thin cover or none at all, are
called _intolerant_. The leaves of tolerant trees endure shade well,
so that their inner and lower leaves flourish under the shadow of
their upper and outer leaves, with the result that the whole tree,
as beech and maple, makes a dense shadow; whereas the leaves of
intolerant trees are either sparse, as in the larch, or are so hung
that the light sifts thru them, as in poplar and oak. The spruces and
balsam fir have the remarkable power of growing slowly under heavy
shade for many years, and then of growing vigorously when the light is
let in by the fall of their overshadowing neighbors. This can plainly
be seen in the cross-section of balsam fir, Fig. 63, where the narrow
annual rings of the early growth, are followed by the wider ones of
later growth. A common sight in the dense woods is the maple sending
up a long, spindly stem thru the trees about it and having at its top
a little tuft of leaves, Fig. 64. By so doing it survives. The fact
that a tree can grow without shade often determines its possession
of a burnt-over tract. The order in the North Woods after a fire
is commonly, first, a growth of fire weed, then raspberries or
blackberries, then aspen, a very intolerant tree whose light shade
in turn permits under it the growth of the spruce, to which it is a
"nurse," Fig. 65. In general it may be said that all seedling conifers
require some shade the first two years, while hardwoods in temperate
climates, as a rule, do not.
[Illustration: Fig. 63. Cross section of Balsam Fir, Showing Fast
Growth After Years of Suppression. Notice the width of the annual
rings in later age compared with early. _U. S. Forest Service._]
[Illustration: Fig. 64. Tolerant Maple. The trees are too slender to
stand alone. _U. S. Forest Service._]
[Illustration: Fig. 65. Intolerant Aspen, a "nurse" of Tolerant
Spruce._ U. S. Forest Service._]
This matter of tolerance has also much to do with the branching of
trees. The leaves on the lower branches of an intolerant tree will not
thrive, with the result that those branches die and later drop off.
This is called "cleaning," or natural pruning. Intolerant trees, like
aspen and tulip, Fig. 66, clean themselves well and hence grow with
long, straight boles, while tolerant trees, like spruce and fir,
retain their branches longer.
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