Cellulose.—I did not obtain this substance in a separate state, but
tried angular bits of dry wood, cork, sphagnum moss, linen, and cotton
thread. None of these bodies were in the least attacked by the
secretion, and they caused only that moderate amount of inflection
which is common to all inorganic objects. Gun-cotton, which consists of
cellulose, with the hydrogen replaced by nitrogen, was tried with the
same result. We have seen that a decoction of cabbage-leaves excites
the most powerful inflection. I therefore placed two little square bits
of the blade of a cabbage-leaf, and four little cubes cut from the
midrib, on six leaves of Drosera. These became well inflected in 12
hrs., and remained so for between two and four days; the bits of
cabbage being bathed all the time by acid secretion. This shows that
some exciting matter, to which I shall presently refer, had been
absorbed; but the angles of the squares and cubes remained as sharp as
ever, proving that the framework of cellulose had not been attacked.
Small square bits of spinach-leaves were tried with the same result;
the glands pouring forth a moderate supply of acid secretion, and the
tentacles remaining inflected for three days. We have also seen that
the delicate coats of pollen grains are not dissolved by the secretion.
It is well known that the gastric juice of animals does not attack
cellulose.
Chlorophyll.—This substance was tried, as it contains nitrogen. Dr.
Moore sent me some preserved in alcohol; it was dried, but soon
deliquesced. Particles were placed on four leaves; after 3 hrs. the
secretion was acid; after 8 hrs. there was a good deal of inflection,
which in 24 hrs. became fairly well marked. After four days two of the
leaves began to open, and the other two were then almost fully
re-expanded. It is therefore clear that this chlorophyll contained
matter which excited the leaves to a moderate degree; but judging by
the eye, little or none was dissolved; so that in a pure state it would
not probably have been attacked by the secretion. Dr. Sanderson tried
that which I [page 126] used, as well as some freshly prepared, with
artificial digestive liquid, and found that it was not digested. Dr.
Lauder Brunton likewise tried some prepared by the process given in the
British Pharmacopoeia, and exposed it for five days at the temperature
of 37° Cent. to digestive liquid, but it was not diminished in bulk,
though the fluid acquired a slightly brown colour. It was also tried
with the glycerine extract of pancreas with a negative result. Nor does
chlorophyll seem affected by the intestinal secretions of various
animals, judging by the colour of their excrement.