The length of time during which the tentacles remain inflected largely
depends on the quantity of the substance given, partly on the facility
with which it is penetrated or acted on by the secretion, and partly on
its essential nature. The tentacles always remain inflected much longer
over large bits or large drops than over small bits or drops. Texture
probably plays a part in determining the extraordinary length of time
during which the tentacles remain inflected [page 133] over the hard
grains of chemically prepared casein. But the tentacles remain
inflected for an equally long time over finely powdered, precipitated
phosphate of lime; phosphorus in this latter case evidently being the
attraction, and animal matter in the case of casein. The leaves remain
long inflected over insects, but it is doubtful how far this is due to
the protection afforded by their chitinous integuments; for animal
matter is soon extracted from insects (probably by exosmose from their
bodies into the dense surrounding secretion), as shown by the prompt
inflection of the leaves. We see the influence of the nature of
different substances in bits of meat, albumen, and fresh gluten acting
very differently from equal-sized bits of gelatine, areolar tissue, and
the fibrous basis of bone. The former cause not only far more prompt
and energetic, but more prolonged, inflection than do the latter. Hence
we are, I think, justified in believing that gelatine, areolar tissue,
and the fibrous basis of bone, would be far less nutritious to Drosera
than such substances as insects, meat, albumen, &c. This is an
interesting conclusion, as it is known that gelatine affords but little
nutriment to animals; and so, probably, would areolar tissue and the
fibrous basis of bone. The chondrin which I used acted more powerfully
than gelatine, but then I do not know that it was pure. It is a more
remarkable fact that fibrin, which belongs to the great class of
Proteids,* including albumen in one of its sub-groups, does not excite
the tentacles in a greater degree, or keep them inflected for a longer
time, than does gelatine, or
* See the classification adopted by Dr. Michael Foster in Watts’
‘Dictionary of Chemistry,’ Supplement 1872, page 969. [page 134]