“Propionic acid 0.0563 Butyric acid 0.0835 Valerianic acid 0.0615
“The quantity digested by a similar liquid containing hydrochloric acid
was 0.3376. Hence, taking this as 100, the following numbers represent
the relative quantities digested by the other acids:—
“Propionic acid 16.5 Butyric acid 24.7 Valerianic acid 16.1
“6. A third experiment of the same kind gave: [page 91]
“Quantity of fibrin digested in four hours by 10 cub. cent. of the
liquid:—
“Hydrochloric acid 0.2915 Propionic acid 0.1490 Butyric acid 0.1044
Valerianic acid 0.0520
“Comparing, as before, the three last numbers with the first taken as
100, the digestive power of propionic acid is represented by 16.8; that
of butyric acid by 35.8; and that of valerianic by 17.8.
“The mean of these three sets of observations (hydrochloric acid being
taken as 100) gives for
“Propionic acid 15.8 Butyric acid 32.0 Valerianic acid 21.4
“7. A further experiment was made to ascertain whether the digestive
activity of butyric acid (which was selected as being apparently the
most efficacious) was relatively greater at ordinary temperatures than
at the temperature of the body. It was found that whereas 10 cub. cent.
of a liquid containing the ordinary proportion of hydrochloric acid
digested 0.1311 gramme, a similar liquid prepared with butyric acid
digested 0.0455 gramme of fibrin.
“Hence, taking the quantities digested with hydrochloric acid at the
temperature of the body as 100, we have the digestive power of
hydrochloric acid at the temperature of 16° to 18° Cent. represented by
44.9; that of butyric acid at the same temperature being 15.6.”
We here see that at the lower of these two temperatures, hydrochloric
acid with pepsin digests, within the same time, rather less than half
the quantity of fibrin compared with what it digests at the higher
temperature; and the power of butyric acid is reduced in the same
proportion under similar conditions and temperatures. We have also seen
that butyric acid, which is much more efficacious than propionic or
valerianic acids, digests with pepsin at the higher temperature less
than a third of the fibrin which is digested at the same temperature by
hydrochloric acid.] [page 92]
I will now give in detail my experiments on the digestive power of the
secretion of Drosera, dividing the substances tried into two series,
namely those which are digested more or less completely, and those
which are not digested. We shall presently see that all these
substances are acted on by the gastric juice of the higher animals in
the same manner. I beg leave to call attention to the experiments under
the head albumen, showing that the secretion loses its power when
neutralised by an alkali, and recovers it when an acid is added.
Substances which are completely or partially digested by the Secretion
of Drosera.