The Foot-prints of the Creator : $b or, The Asterolepis of Stromness
John Stuart Mill · en
Agassiz ultimately succeeded in classing the bones which had at first so
puzzled him, into two divisions—interior and dermal; and the latter he
divided yet further, though not without first lodging a precautionary
protest, founded on the extreme obscurity of the subject, into cranial
and opercular. Of the interior bones he specified two,—a super-scapular
bone, (_supra-scapulaire_,)—that bone which in osseous fishes completes
the scapular arch or belt, by uniting the scapula to the cranium; and
a maxillary or upper jaw-bone. But his world-wide acquaintance with
existing fishes could lend him no assistance in determining the places
of the dermal bones: they formed the mere fragments of a broken puzzle,
of which the key was lost. Even in their detached and irreducible state,
however, he succeeded in basing upon them several shrewd deductions. He
inferred, in the first place, that the _Asterolepis_ was not, as had
been at first supposed, a cuirassed fish, which took its place among
the Cephalaspians, but a strongly helmed fish of that Cœlacanth family
to which the _Holoptychius_ and _Glyptolepis_ belong; in the second,
that, like several of its bulkier cogeners, it was in all probability
a broad, flat-headed animal; and, in the third, that as its remains
are found associated in the Russian beds with numerous detached teeth
of large size,—the boar tusks of Kutorga—which present internally that
peculiar microscopic character on which Professor Owen has erected his
Dendrodic or tree-toothed family of fishes,—it would in all likelihood
be found that both bones and teeth belonged to the same group. “It
appears more than probable,” he said, “that one day, by the discovery of
a head or an entire jaw, it will be shown that the genera _Dendrodus_
and _Asterolepis_ form but one.” As we proceed, the reader will see how
justly the ichthyologist assigned to the _Asterolepis_ its place among
the Cœlacanths, and how entirely his two other conjectures regarding it
have been confirmed. “I have had in general,” he concluded, “but small
and mutilated fragments of the creature’s bones submitted to me, and
of these, even the surface ornaments not well preserved; but I hope
the immense materials with which the Old Red Sandstone of Russia has
furnished the savans of that country will not be lost to science; and
that my labors on this interesting genus, incomplete as they are, will
excite more and more the attention of geologists, by showing them how
ignorant we are of all the essential facts concerning the history of the
first inhabitants of our globe.”