The Foot-prints of the Creator : $b or, The Asterolepis of Stromness
John Stuart Mill · en
The scales of the Cœlacanths were, in almost all the genera which
compose the family, of great size; in some species, of the greatest size
to which this kind of integument ever attained. Of a Cœlacanth of the
Coal Measures, the _Holoptychius Hibberti_, the scales in the larger
specimens were occasionally from five to six inches in diameter. Even
in the _Holoptychius nobilissimus_, in an individual scarcely exceeding
two and a half feet in length, they measured from an inch and a half to
an inch and three quarters each way. In the splendid specimen of this
last species, in the British Museum, there occur but fourteen scales
between the ventrals, though these lie low on the creature’s body, and
the head; and in a specimen of a smaller species,—the _Holoptychius
Andersoni_,—but about seventeen. The exposed portion of the scale was
in most species of the family curiously fretted by intermingled ridges
and furrows, pits and tubercles, which were either boldly relieved, as
in the _Holoptychius_, or existed, as in the _Glyptolepis_, as slim,
delicately chiselled threads, lines, and dots. The head was covered by
strong plates, which were roughened with tubercles either confluent or
detached, or hollowed, as in the _Bothriolepis_, into shallow pits. The
jaws were thickly set with an outer range of true fish teeth, and more
thinly with an inner range of what seem _reptile_ teeth, that stood up,
tall and bulky, behind the others, like officers on horseback seen over
the heads of their foot-soldiers in front. The _double_ fins,—pectorals
and ventrals,—were characterized each by a thick, angular, scale-covered
centre, fringed by the rays; and they must have borne externally
somewhat the form of the sweeping paddles of the Ichthyosaurian genus,—a
peculiarity shared also by the double fins of the _Dipterus_. The
_single_ fins, in all the members of the family of which specimens
have been found sufficiently entire to indicate the fact, were four in
number,—an anal, a caudal, and two dorsal fins; and, with the exception
of the anterior dorsal, which was comparatively small, and bent downwards
along the back, as if its rays had been distorted when young,[11] they
were all of large size. They crowded thickly on the posterior portion of
the body,—the anterior dorsal opposite the ventrals, and the posterior
dorsal opposite the anal fin. The fin-rays of the various members of the
family, and such of their spinous processes as have been detected, were
hollow tubular bones; or rather, like the larger pieces in the framework
of the Placoids, they were cartilaginous within, and covered externally
by a thin osseous crust or shell, which alone survives; and to this
peculiarity they owe their family name, Cœlacanth, or “hollow-spine.”
The internal hollow, _i. e._ cartilaginous centre, was, however,
equally a characteristic of the spinous processes of the _Coccosteus_.
In their general proportions, the Cœlacanths, if we perhaps except one