After self-thinning has gone on for a hundred years or so, the ranks
have been so thinned that there are openings sufficiently large to
allow other species a chance to come in. By this time, too, there is
sufficient humus on the floor to allow the seeds of many other species
to germinate. Lodge-pole thus colonizes barren places, holds them for
a time, and so changes them that the very species dispossessed by fire
may regain the lost territory. Roughly, the lodge-pole will hold the
ground exclusively from seventy-five to one hundred and fifty years,
then the invading trees will come triumphantly in and, during the next
century and a half, will so increase and multiply that they will
almost exclude the lodge-pole. Thus Engelmann spruce and Douglas fir
are now growing where lodge-pole flourished, but let fire destroy this
forest and lodge-pole will again claim the territory, hold it against
all comers for a century or two, and then slowly give way to or be
displaced by the spruces and firs.
The interesting characteristic of holding its cones and hoarding seeds
often results in the cones being overgrown and embedded in the trunk
or the limbs of the trees. As the cones hug closely the trunk or the
limbs, it is not uncommon for the saw, when laying open a log at the
mill, to reveal a number of cones embedded there. I have in my cabin a
sixteen-foot plank that is two inches in diameter and six inches wide,
which came out of a lodge-pole tree. Embedded in this are more than a
score of cones. Probably most of these cones were of the first crop
which the tree produced, for they clung along the trunk of the tree
and grew there when it was about an inch and a quarter in diameter.
The section upon which these cones grew was between fifteen and
twenty-five feet from the ground.
The seeds of most conifers need vegetable mould, litter, or vegetation
cover of some kind in which to germinate, and then shade for a time in
which to grow. These requirements so needed by other conifer seeds and
seedlings are detrimental to the lodge-pole. If its seeds fall on
areas lightly covered with low huckleberry vines, but few of them will
germinate. A lodge-pole seed that germinates in the shade is doomed.
It must have sunlight or die. In the ashes of a forest fire, in the
full glare of the sun, the seeds of the lodge-pole germinate, grow,
and flourish.
Wind is the chief agency which enables the seeds to migrate. The seeds
are light, and I know of one instance where an isolated tree on a
plateau managed to scatter its seeds by the aid of the wind over a
circular area fifty acres in extent, though a few acres is all that
is reached by the average tree. Sometimes the wind scatters the seeds
unevenly. If most of the seeds are released in one day, and the wind
this day prevails from the same quarter, the seeds will take but one
course from the tree; while changing winds may scatter them quite
evenly all around the tree.