A Critique of the Theory of EvolutionMorgan, Thomas Hunt
Science
A Critique of the Theory of Evolution
Morgan, Thomas Hunt
Evolution
In the preceding cases I have spoken as though a factor influenced only one
part of the body. It would have been more accurate to have stated that the
_chief_ effect of the factor was observed in a particular part of the body.
Most students of genetics realize that a factor difference usually affects
more than a single character. For example, a mutant stock called
rudimentary wings has as its principle characteristic very short wings
(fig. 34). But the factor for rudimentary wings also produces other effects
as well. The females are almost completely sterile, while the males are
fertile. The viability of the stock is poor. When flies with rudimentary
wings are put into competition with wild flies relatively few of the
rudimentary flies come through, especially if the culture is crowded. The
hind legs are also shortened. All of these effects are the results of a
single factor-difference.
[Illustration: FIG. 34. Mutant race of fruit fly, called rudimentary.]
One may venture the guess that some of the specific and varietal
differences that are characteristic of wild types and which at the same
time appear to have no survival value, are only by-products of factors
whose most important effect is on another part of the organism where their
influence is of vital importance.
It is well known that systematists make use of characters that are constant
for groups of species, but which do not appear in themselves to have an
adaptive significance. If we may suppose that the constancy of such
characters may be only an index of the presence of a factor whose _chief_
influence is in some other direction or directions, some physiological
influence, for example, we can give at least a reasonable explanation of
the constancy of such characters.
I am inclined to think that an overstatement to the effect that each factor
may affect the entire body, is less likely to do harm than to state that
each factor affects only a particular character. The reckless use of the
phrase "unit character" has done much to mislead the uninitiated as to the
effects that a single change in the germ plasm may produce on the organism.
Fortunately, the expression "unit character" is being less used by those
students of genetics who are more careful in regard to the implications of
their terminology.
There is a class of cases of inheritance, due to the XY chromosomes, that
is called sex linked inheritance. It is shown both by mutant characters and
characters of wild species.
For instance, white eye color in Drosophila shows sex linked inheritance.
If a white eyed male is mated to a wild red eyed female (fig. 35) all the
offspring have red eyes. If these are inbred, there are three red to one
white eyed offspring, but white eyes occur only in the males. The
grandfather has transmitted his peculiarity to half of his grandsons, but
to none of his granddaughters.
Public-domain text, read in full here on John Shaqi.
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