Produced by Chris Curnow, Joseph Cooper, Diane Monico, and the Online Distributed Proofreading Team at http://www.pgdp.net UNIVERSITY OF KANSAS PUBLICATIONS MUSEUM OF NATURAL HISTORY Volume 13, No. 2, pp. 19-72, pls. 1-8, 3 figs. August 16, 1960 A Distributional Study of the Amphibians of the Isthmus of Tehuantepec, Mexico BY WILLIAM E. DUELLMAN UNIVERSITY OF KANSAS LAWRENCE 1960 UNIVERSITY OF KANSAS PUBLICATIONS MUSEUM OF NATURAL HISTORY Volume 13, No. 2, pp. 19-72, pls. 1-8, 3 figs. August 16, 1960 A Distributional Study of the Amphibians of the Isthmus of Tehuantepec, Mexico BY WILLIAM E. DUELLMAN UNIVERSITY OF KANSAS LAWRENCE 1960 UNIVERSITY OF KANSAS PUBLICATIONS, MUSEUM OF NATURAL HISTORY Editors: E. Raymond Hall, Chairman, Henry S. Fitch, Robert W. Wilson Volume 13, No. 2, pp. 19-72, pls. 1-8, 3 figs. Published August 16, 1960 UNIVERSITY OF KANSAS Lawrence, Kansas PRINTED IN THE STATE PRINTING PLANT TOPEKA, KANSAS 1960 28-3859 A Distributional Study of the Amphibians of the Isthmus of Tehuantepec, Mexico BY WILLIAM E. DUELLMAN CONTENTS PAGE INTRODUCTION 21 Acknowledgments 23 Field Studies in the Isthmus of Tehuantepec 23 Sources of Material 24 DESCRIPTION OF THE ISTHMUS OF TEHUANTEPEC 25 Physiography 25 Climate 28 Vegetation 29 The Sierra de los Tuxtlas 32 GAZETTEER 33 THE AMPHIBIAN FAUNA OF THE LOWLANDS 37 Composition of the Fauna 37 Ecology of the Fauna 38 Distribution of the Fauna 42 THE AMPHIBIAN FAUNA OF THE FOOTHILLS AND ADJACENT HIGHLANDS 44 ESTABLISHMENT OF PRESENT PATTERNS OF DISTRIBUTION 45 ACCOUNTS OF SPECIES 49 SUMMARY 68 LITERATURE CITED 69 INTRODUCTION