A first book in organic evolutionShute, D. Kerfoot (Daniel Kerfoot)
Science
A first book in organic evolution
Shute, D. Kerfoot (Daniel Kerfoot)
Evolution
The equatorial plate of split V-shaped
mother chromosomes (hereditary threads) thus divides the fibers of the
spindle into two parts, one half extending from one centrosome to one
group of daughter chromosomes, while the remaining half extend from the
other centrosome to the other group of daughter chromosomes. Soon the
achromatin fibers of the spindle contract, and in this way separate the
two groups of daughter chromosomes, so that one group is drawn towards
one centrosome, and the other group to the other centrosome (Fig. 8, E,
F, G, H). After the two groups of daughter chromosomes have been drawn to
their respective centrosomes, each group assumes the tangle or network
phase like the nucleus of the mother cell, and an investing nuclear
membrane reappears for each (Fig. 8, I). Thus from the mother nucleus of
the mother cell we get two daughter nuclei (I). In a further phase of
the mitotic process, a furrow appears on the surface of the protoplasm
and surrounds it in the form of a ring. This furrow is in a plane at
right angles to the long axis of the spindle, and gradually deepens until
the protoplasm is divided into two parts, each segment of protoplasm
containing its own nucleus and centrosome; in short, the mother cell has
divided into two daughter cells (I).
It will thus be observed that the centrosomes and their achromatin fibers
are a beautiful mechanism by which the heredity threads (chromosomes) are
exactly divided into two equivalent halves.
There are some cells (_Amœba proteus_, for instance,) which divide in a
much simpler manner than by mitosis; in these there is no complicated
rearrangement of the chromosomes and no disappearance of the nuclear
membrane, the nucleus simply becoming separated into two parts
(_Amitosis_).
=The Human Ovum.= The human ovum is a typical cell about one-fifth
of a millimeter in size and spherical in shape. It is a nucleated
piece of protoplasm possessing an enveloping cell-wall (metaplasm).
The protoplasm contains nutrient material or yolk (metaplasm). The
_maturation_ of the ovum essentially consists in throwing out half of its
chromosomes. In doing this the nucleus (mother) approaches the surface of
the protoplasm (Fig. 9, A), and divides, by mitosis, into two daughter
nuclei; then the unripe ovum divides into two cells, but of very unequal
size. This process is repeated a second time. Thus two small cells are
formed which are known as the _polar bodies_ (Fig. 9, A, B, pol. b). The
large cell remaining after the formation of the two polar bodies is the
_mature ovum_ (B). Its nucleus, which recedes towards the center of the
protoplasm, is called the female _pronucleus_ (Fig. 9, B, f. pr.). The
pronuclei of mature ova differ from the nuclei of all the other cells of
the body in that they only contain half as many chromosomes (hereditary
threads).
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