A Guide to the Study of Fishes, Volume 2 (of 2)Jordan, David Starr
History
A Guide to the Study of Fishes, Volume 2 (of 2)
Jordan, David Starr
Fishes; Fishes -- Bibliography; Zoology -- History
body may be found among the Ganoids. In the Mesozoic seas these fishes
were scarcely less varied and perhaps scarcely less abundant than the
Teleosts in the seas of to-day. They far exceed the Crossopterygians in
number and variety of forms. Transitional forms connecting the two
groups are thus far not recognized. So far as fossils show, the
characteristic actinopterous fin with its reduced and altered basal
bones appeared at once without intervening gradations.
The name _Ganoidei_ (γάνος, brightness; εἶδος, resemblance), alluding to
the enameled plates, was first given by Agassiz to those forms, mostly
extinct, which were covered with bony scales or hard plates of one sort
or another. As the term was originally defined, mailed catfishes,
sea-horses, _Agonidæ_, _Arthrodires_, _Ostracophores_, and other wholly
unrelated types were included with the garpikes and sturgeons as
Ganoids. Most of these intruding forms among living fishes were
eliminated by Johannes Müller, who recognized the various archaic
characters common to the existing forms after the removal of the mailed
Teleosts. Still later Huxley separated the Crossopterygians as a
distinct group, while others have shown that the _Ostracophori_ and
_Arthrodira_ should be placed far from the garpike in systematic
classification. Cope, Woodward, Hay, and others have dropped the name
Ganoid altogether as productive of confusion through the many meanings
attached to it. Others have kept it as a convenient group name for the
orders of archaic _Actinopteri_. For these varied and more or less
divergent forms it seems convenient to retain it. As an adjective
"ganoid" is sometimes used as descriptive of bony plates or enameled
scales, some-in the sense of archaic, as applied to fishes.
=Are the Ganoids a Natural Group?=—Several writers have urged that the
_Ganoidei_, even as thus restricted, should not be considered as a
natural group, whether subclass, order, or group of orders. The reasons
for this view in brief are the following:
1. The group is heterogeneous. The _Amiidæ_ differ more from the other
Ganoids than they do from the herring-like Teleosts. The garpikes,
sturgeons, paddle-fishes likewise diverge widely from each other and
from the _Palæoniscidæ_ and the _Platysomidæ_. Each of the living
families represents the residue or culmination of a long series, in some
cases advancing, as in the case of the bowfin, sometimes perhaps
degenerating, as in the case of the sturgeons.
2. Of the traits possessed in common by these forms, several (the
cellular air-bladder, the many valves in the heart, the spiral valve in
the intestine, the heterocercal tail) are all possessed in greater or
less degree by certain _Isospondyli_ or allies of the herring. All these
characters are still better developed in _Crossoptergyii_ and
_Dipneusti_, and each one disappears by degrees. Of the characters drawn
from the soft parts we can know nothing so far as the extinct Ganoids
are concerned.
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