A Handbook of Some South Indian GrassesRangachari, K.
Science
A Handbook of Some South Indian Grasses
Rangachari, K.
Grasses -- India, South; Plants -- India, South
The stem of _Panicum fluitans_ is round in outline in the transverse
section and has a large cavity. Just close to the cavity and separated
from it by only one or two parenchymatous cells are found vascular
bundles forming a series. Outside this series of bundles lies a
sclerenchymatous band which is wavy, following the lower edges of the
large air-cavities. One series of bundles is connected with this
sclerenchymatous ring. The air-cavities are large and uniform and are
separated by bands of parenchymatous cells. In each of these bands lies
a vascular bundle on the upper side near the periphery. Sometimes we
find, especially in young stages, diaphragms of stellulate cells
stretched across the air-cavities. Later as the stem matures these
disappear and the cavities become conspicuous. (See figs. 42 and 43.)
=Structure of the root.=--As already stated, the roots of grasses
conform to the monocotyledonous type, but the variations met with in
their structure are not so great as in the case of the stem. The
root-tips are protected by root-caps, and the actual tip of the root is
very distinct in the roots of all grasses and it can be seen very
clearly in a longitudinal section of the root. The actual tip of the
root is sharply distinct from the root-cap as there are two distinct
sets of cells, one giving rise to the root-tip and the other to the
root-cap.
The young root-tips are always free from root-hairs, and they are
confined to the portions behind the root-tips. The extent of the
root-hair region will vary according to the vigour and development of
the roots and the nature of the soil. The root-hairs are mere
protrusions of the cells of the outermost layer of the cortex of the
root and this layer is called the =piliferous layer=.
[Illustration: Fig. 44.--Transverse section of a part of the root of
Pennisetum cenchroides. x 100
1. Root-hair; 2. piliferous layer; 3. cortex; 4. endodermis; 5.
pericycle; 6. xylem; 7. phloem.]
To learn the structure of the roots of grasses we may select as types
the roots of _Pennisetum cenchroides_ and _Andropogon Sorghum_ and
consider their structural details. In the transverse sections of these
roots we find a fairly broad cortex consisting of thin-walled
parenchymatous cells more or less regularly arranged. (See figs. 44 and
45.) Just below the piliferous layer two or three layers of thick-walled
cells are seen. In the roots of _Andropogon Sorghum_ these thick-walled
cells are very conspicuous as they consist of several layers. These
layers of thick-walled cells constitute the =exodermis=. (See fig. 46.)
The innermost layer of cells of the cortex is called the =endodermis=
and it becomes conspicuous on account of the thickening in the lateral
and inner walls of the cells of this layer. (See figs. 44 and 47.)
[Illustration: Fig. 45.--Transverse section of the entire root of
Andropogon Sorghum. x 25]
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