A Handbook of Some South Indian GrassesRangachari, K.
Science
A Handbook of Some South Indian Grasses
Rangachari, K.
Grasses -- India, South; Plants -- India, South
The sclerenchyma lying on the lower side of the primary bundles are
contiguous with the bundle, while those above are separated from the
bundle by the chlorophyllous layer. (See fig. 55.) In the case of
secondary and tertiary bundles the sclerenchymatous bands lying on the
lower side are in contact with the chlorophyllous layer, whereas the
upper bands are either in contact with this layer or separated from it
by a few parenchymatous cells.
All the vascular bundles in the leaves of _Eragrostis Willdenoviana_ are
provided with sclerenchyma on both the sides. The lower band of the
primary vascular bundles is continuous with the vascular bundle, the
chlorophyllous layer being open below. The upper bands of the primary
and the lower bands of the secondary vascular bundles just touch the
chlorophyllous layer. In the secondary bundles the sclerenchyma band
above is separated from the chlorophyllous layer by two layers of
parenchyma. In the case of the leaves of _Panicum flavidum_, _P.
colonum_, _P. fluitans_ and _Pennisetum cenchroides_ the sclerenchyma is
separated from the chlorophyllous layer by layers of parenchyma.
[Illustration: Fig. 56.--Transverse section of a part of the leaf of
Eragrostis Willdenoviana. x 30
1. Vascular bundle; 2. sclerenchyma; 3. motor cells.]
[Illustration: Fig. 57.--Transverse section of a portion of the leaf of
Eragrostis Willdenoviana. x 150
1. Motor cells; 2. stomata; 3. sclerenchyma; 4. chlorophyllous layer; 5.
vascular strand cut through; 6. hair.]
[Illustration: Fig. 58.--Transverse section of a part of the leaf of
Panicum colonum. x 30
1. Vascular bundle; 2. sclerenchyma; 3. motor cells.]
Even from the few examples dealt with above, it is obvious that the
range of variation of sclerenchyma in leaves is very great. In the
leaves of _Aristida setacea_ there is a considerable amount of
sclerenchyma whilst in some leaves such as those of _Panicum_ _colonum_,
_P. flavidum_ and _Panicum fluitans_ the sclerenchyma is reduced to its
minimum.
[Illustration: Fig. 59.--Transverse section of a part of the leaf of
Panicum fluitans. x 30
1. Vascular bundle; 2. sclerenchyma.]
In the leaves of grasses growing in dry situations the development of
sclerenchyma is generally very considerable. The grass _Aristida
setacea_ is a good example of a xerophytic grass. The sea-shore grass
_Spinifex squarrosus_ is another example of the same kind. But in the
leaves of this grass, the development of sclerenchyma is not very
considerable, but there is a great development of parenchymatous cells
free from chlorophyll within the leaf, the chlorophyll bearing cells
being confined to the upper and the lower surfaces of the leaves.
[Illustration: Fig. 60.--Transverse section of a leaf of Spinifex
squarrosus. x 10
1. Vascular bundle; 2. sclerenchyma.]
[Illustration: Fig. 61.--A portion of the transverse section of the leaf
of Spinifex squarrosus. x 60
1. Sclerenchyma; 2. vascular bundle; 3. epidermis; 4. stomata.]
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