A Handbook of Some South Indian GrassesRangachari, K.
Science
A Handbook of Some South Indian Grasses
Rangachari, K.
Grasses -- India, South; Plants -- India, South
As regards size, the epidermal cells overlying the sclerenchyma are
small and those lying over parenchyma are larger. Amongst the larger
cells some may be motor-cells. The stomata occur in regular rows between
the vascular bundles and they are quite characteristic of grasses. They
are more or less similar in structure in all grasses. In the leaves of
many grasses stomata are found in both the upper and the lower epidermis
and they are confined to the lower epidermis in a few grasses only.
The motor-cells vary very much both as regards their shape and position.
In some leaves as in the leaves of the grass _Panicum flavidum_ the
motor-cells are confined to the midrib on the upper surface.
The epidermal cells of this leaf are large and uniformly round. (See
figs. 66 and 67.)
[Illustration: Fig. 66.--Transverse section of a leaf of Panicum
flavidum. x 20
1. Vascular bundle; 2. sclerenchyma; 3. motor-cells.]
[Illustration: Fig. 67.--Transverse section of the leaf of Panicum
flavidum. x 150
1. Motor-cells; 2. stomata; 3. sclerenchyma; 4. chlorophyllous layer.]
In the case of most grasses the motor-cells are found in groups of
three, four or five between the vascular bundles. The central motor-cell
is usually the largest and it is somewhat obovate in shape in a
transverse section of the leaf. In the leaves of _Panicum javanicum_ and
_Eriochloa polystachya_ there are three or four motor cells in the group
and the group consists of four, five or rarely six motor cells in the
leaves of _Eragrostis Willdenoviana_. When there are distinct furrows
between ridges these cells lie in the furrows and they are many in
number. In the leaves of _Panicum repens_ there are five to seven
motor-cells in the furrows and the single row of cells stretched between
the motor-cells and the lower epidermis in the furrow consists of more
or less clear cells with sparsely scattered small chlorophyll grains.
(See fig. 52.) The motor-cells occupying the furrows in the leaves of
_Aristida setacea_ are more in number than in _Panicum repens_ and are
of a different shape. All the cells lying in the furrow between the
motor-cells and the sclerenchyma are clear cells free from chlorophyll
grains.
Although the motor-cells differ in shape from the ordinary epidermal
cells in most grasses, there are, however, a few grasses in which the
motor-cells do not differ very much from the epidermal cells except in
size. For example, in the leaves of _Panicum colonum_ the motor-cells
are just like the ordinary epidermal cells in shape but are larger. (See
fig. 64.)
Motor-cells are usually confined to the upper epidermis, but they may
also be found in the lower epidermis. In the leaves of _Pennisetum
cenchroides_ motor-cells are found in both the upper and the lower
epidermis, the group in the upper epidermis alternating with that in the
lower.
[Illustration: Fig. 68.--Transverse section of a portion of the leaf of
Pennisetum cenchroides. x 100
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