In many species the ascospores germinate and form conidia
immediately (_Nectria cinnabarina_, _Sclerotinia_, _Taphrina_,
etc.), sometimes while they are still in the ascus and before
their ejection (_Taphrina_, Fig. 105 _a_). In many instances
the conidia by means of continued budding can, for a longer
or shorter time, produce yeast-conidia, _e.g. Taphrina_. In
many other cases the conidia arise from the germ-tubes of the
ascospores, or at any part of the mycelium. The unripe asci
of _Taphrina_, when placed in water, develop conidia at their
apices. The _Sclerotinia_-species produce numerous conidia
whose germination has never been observed. The formation of
conidia and asci sometimes takes place on the same fruit-body.
In _Heterosphæria patella_ the conidia and asci are developed
successively in the same fruit-body; in the ascocarps of
_Dermatea frangula_ and _Sclerotinia sclerotiorum_ the formation
of conidia may take place. The ascocarps frequently arise
from the conidial-layers (_Nectria cinnabarina_, etc.). This
relationship of the two forms of reproduction to each other
may be explained by considering that both have descended
phylogenetically from sporangia.
Sometimes _chlamydospores_ and _oidia_ also appear in the Ascomycetes;
on germination, however, they do not, as in _Protomyces_, form
sporangia, and on this account cannot be distinctly distinguished from
conidia.
The asci are morphologically the highest form of reproduction and
are always found at the close of the development of these Fungi; the
accessory forms of reproduction are first developed, but a well-defined
alternation of generations does not occur.
In the Ascomycetes there are more than 11,000 described species,
which can be classed as follows:--
Series 1. EXOASCI. Only one order.
„ 2. CARPOASCI.
Family 1. _Gymnoascales_, }
„ 2. _Perisporiales_, } Angiocarpic Carpoasci.
„ 3. _Pyrenomycetes_, }
„ 4. _Hysteriales_, } Hemiangiocarpic Carpoasci.
„ 5. _Discomycetes_,}
„ 6. _Helvellales_, Gymnocarpic (?) Carpoasci.
Additional _Ascolichenes_: Lichen-forming Ascomycetes.
Series 1. =Exoasci.=
Ascomycetes with FREE ASCI; sometimes also conidia, chlamydospores and
oidia. One order.
Order. =Taphrinaceæ.= Of the genera belonging to this order,
_Taphrina_, _Endomyces_, and _Ascocorticium_, the first is most
important.
_Endomyces decipiens_ is a parasite in the fruit-body of
_Armillaria mellea_; _E. magnusii_ lives in the gelatinous,
fermenting exudations of Oak-trees; _Ascocorticium albidum_
is found under the bark of the Fir-tree. _Endomyces_ has
chlamydospores and oidia.
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