The MYCOMYCETES are not entirely aquatic in habit; they have hyphæ with
_transverse walls_, but _no sexual reproductive organs_. The asexual
reproduction takes place in very different ways; by endospores (in
asci), conidia, basidiospores, chlamydospores, and oidia. Swarmspores
are never found.
Two chief methods of reproduction may be distinguished, and hence the
class may be divided into two large sub-classes:--the ASCOMYCETES (with
asci), and the BASIDIOMYCETES (with basidia).
Sub-Class 1. =Ascomycetes.=
The main characteristic which distinguishes the Ascomycetes is the
_ascus_; a name given to a sporangium of a definite shape and size,
and containing a definite number of spores. The shape is generally
club-like or spherical, the number of spores 8 (in some 2, 4, 16 or
more), see Figs. 103, 105, 108, 110, 113, 116, 120, 121, 123, 129.
In the lowest forms, the EXOASCI, the ascus springs directly from the
mycelium without the formation of a fruit-body (_i.e._ ascocarp).
In the higher forms, which contain many species, the CARPOASCI, the
asci are united and form ascocarps which may be more or less enclosed
(angiocarpic, hemiangiocarpic, and probably gymnocarpic).
[Illustration: FIG. 103.--Endogenous formation of spores in
_Peziza confluens_. In the youngest asci there is only one
nucleus (_b_, _e_); this divides into two (_f_); and the division
is repeated so that there are 4 nuclei in _c_ and 8 in _g_. These
surround themselves with protoplasm and a cell-wall (_h_, _i_).
The protoplasm of the mother-cell is not entirely used up.]
The hyphæ of the _Mycelium_ in some remain free, in others they are
felted together and form thick strands or flat, cushion-like bodies
(compare in particular the stromata of the Pyrenomycetes). Some species
form _sclerotia_ (Figs. 116, 128).
Asexual reproduction by means of _conidia_ is known in many species
as the principal means of reproduction, and the one which affords
the most rapid means of distribution. The conidia may be produced on
conidiophores (Fig. 109), in conidial-layers (Fig. 122), and often in
conidiocarps (pycnidia, Figs. 120 _d_, _e_; 123 _a_; 124 _b._). These
last occur partly as the so-called “spermogonia” (that is, pycnidia
with microconidia). The conidiophores never approach the basidia.
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