The ~FERTILISED OOSPHERE~ divides into an upper (hypobasal) and a lower
(epibasal) cell; from the latter alone the embryo is developed with
its root, stem, foot, and two _cotyledons_, and the former gives rise
to an organ which appears in this instance for the first time, but
which occurs in all Flowering-plants, viz. the _suspensor_. This forces
the embryo down into the “endosperm,” which is entirely or partially
absorbed by the embryo. In the case of the Flowering-plants the embryo
is developed with its longitudinal axis in the elongation of the
suspensor, but in _Selaginella_ the embryo is situated _transversely_
to it.
_Selaginella_ (300–400 species), is essentially tropical, only one
species living in the North (_S. spinulosa_), but others grow in
Central and South Europe.
Order 3. =Lepidodendraceæ= are extinct, tree-like Lycopods,
which are found especially in the Lower and Middle
Carboniferous. Vegetatively they are most nearly related to
_Lycopodium_, but the stem attained much larger proportions
(about eleven metres in height and one metre in thickness),
and had a cambium by which it increased in thickness. It was
regularly dichotomous, and closely studded with spirally-placed
leaves, which left behind them peculiar rhombic scars. The large
cones resemble Pine-cones, and bore sporangia much larger than
any which are now produced (the male ones as much as 2 cm.’s in
length). The macrosporangia were situated at the base, and the
microsporangia at the apex.
Order 4. =Sigillariaceæ.= These are, presumably, another
group of extinct tree-like Lycopods (especially in the Middle
Carboniferous). The name has been derived from the seal-like
scars, which the fallen leaves have left behind in longitudinal
rows on the grooved stem. The rhizomes of these plants were
formerly termed _Stigmaria_, and placed in a separate genus.
Order 5. =Sphenophyllaceæ= form an entirely extinct group.
They do not definitely belong to any of the three large
classes of Vascular Cryptogams, but it is perhaps best to
place them in juxtaposition to these. They were herbaceous
plants with verticillate, wedge-shaped leaves, with nerves
branching dichotomously into equally strong branches. Micro-and
macrosporangia were formed in the same cone; and were situated
in the axils of the leaves, as in the Lycopods.
The Transition from the Cryptogams to the Phanerogams.
Public-domain text, read in full here on John Shaqi.
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