All the plants considered in the preceding chapters are included
in the term CRYPTOGAMS; all in the following chapters under the
head of PHANEROGAMS (see page 3). Hofmeister’s pioneer works
(1851, _Vergleichende Untersuchungen der höheren Kryptogamen_,
etc.) and the numerous researches published later by other
investigators, have closed the gap which was formerly thought
to exist between these plants; so that we now, in the series:
Bryophyta--Pteridophyta--Gymnospermæ--Angiospermæ see the expression of
a single line of development in accordance with a definite plan. The
forms through which this gradual development has taken place have in
course of time, however, to a great extent died out, and only single
links of the chain connecting the lowest to the highest still remain.
THE ALTERNATION OF GENERATIONS, which we found indicated in certain
Thallophytes, can be proved with the greatest clearness in all the
higher Cryptogams, from the Mosses upwards; it is also found in the
Phanerogams, but not in such a pronounced degree, because one of
the generations is so far reduced that it has almost given up its
independence. For the sake of greater clearness, we will begin with the
comparison of the sporophyte, asexual (second) generation.
=The asexual (2nd) generation of the Cormophytes.=
The asexual generation which follows from the further development of
the fertilised oosphere, is, in the _Mosses_, only the sporogonium
(according to one theory it is perhaps homologous with a spore-bearing
leaf, situated upon a short stem, see p. 187); in _Filicinæ_,
_Equisetinæ_, and _Lycopodinæ_, on the other hand, it is a highly
developed plant differentiated into stem, leaf, and true root, and
bearing the sporangia on its leaves. The ~MODIFICATION OF THE SHOOT~
is very slight in _Filicinæ_. The first leaves of the embryo are very
simple in form (Fig. 205), but after a certain age all the leaves
which arise are essentially alike. The fertile leaves do not differ
from the barren ones, and are found associated with them, and their
formation does not limit the growth in length of the stem. It is only
in a few of the true Ferns, and in the Hydropterideæ, that the fertile
leaves differ considerably from the barren ones. A division of labour
in which certain leaves are set apart for nutrition, and others for
reproduction, is found more pronouncedly in the _Equisetinæ_ and
_Lycopodinæ_, for in these groups, with a few exceptions, the fertile
and barren leaves are very dissimilar; the former are collected in
special ear-like _cones_, which _terminate the further growth_ of
the short stems on which they are borne. In connection with the cone,
leaves are sometimes developed which form a transition from the barren
to the fertile ones (the “annulus” in Equisetaceæ), and in these cases
the first indication of a flower with perianth or floral-leaves is to
be traced. Among the Cryptogams the highest division of labour is found
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