in _Selaginella_ and _Isoëtes_, which have the two kinds of sporangia
borne on _different_ leaves. The division of labour (modification) is,
however, still more pronounced in the _Phanerogams_: the leaves which
bear the microsporangia (“pollen-sacs”) have quite different forms
from those which bear macrosporangia (the “nucellus” in the ovule),
the former are termed _stamens_, the latter _carpels_; in certain
instances, too, there is even a contrast between the “male plants” and
the “female plants.” Moreover, a division of labour, in a much greater
degree, takes place in the leaves which do not directly take part in
reproduction, and it is thus possible in many plants to draw a sharp
line not only between stamens and carpels, but also between four or
five distinct kinds of leaves, which differ in _form_, _structure_, and
corresponding _functions_, and which appear in regular sequence on the
shoot: namely, between “scale-leaves” and “foliage-leaves,”[21] both
of which occur in the Cryptogams, and the “floral-leaves,” including
the bracts and leaves of the “perianth,” which latter often differ from
each other in form and colour, and are then separated into _sepals_ and
_petals_. The _leaves_--stamens and carpels--_which bear the sporangia_
are termed sporophylls, and the shoot, or extremity of a shoot, whose
leaves are modified into sporophylls, is _terminated in its further
growth by their production, and is known as a flower_. The flower
which is most completely furnished has calyx, corolla, stamens, and
carpels arranged in this order. If the only sporophylls present are
stamens, then it is said to be a _male_ (_staminate_) flower, and if
only carpels, then a _female_ (_pistillate_) flower, and in both these
cases the flowers are _unisexual_, or diclinous. If stamens and carpels
are both present in the same flower, it is termed _hermaphrodite_.
Diclinous plants in which the female flowers are situated on one plant,
and the male flowers on another, are termed _diœcious_; and those
in which the same plant bears the two kinds of flowers are termed
_monœcious_. When the male, female, and hermaphrodite flowers are found
in the same species, the plant is said to be _polygamous_.
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