_Abies_ (Fir). The leaves are often (_e.g. Ab. pectinata_) displaced
into 2 rows, flat and indented at the apex, with 2 white (wax-covered)
lines on the under surface, in which the stomata are situated. The
leaf-scars are nearly circular and do not project. The cones are erect.
_The cover-scales and the ovuliferous scales separate from the axis_,
to which they remain attached in other genera.--~_Tsuga_ has leaves
like _Abies_, but by the slightly projecting leaf-scars, and cones with
persistent scales, it forms the transition to _Picea_.--_Pseudotsuga_
has leaves similar to those of _Abies_ and persistent carpels as in
_Picea_, but the cover-scales grow as in _Abies_ and project beyond
the ovuliferous scales (_P. douglasii_, Fig. 267). These two genera
are considered as sub-genera of _Abies_.~--_Picea_. The leaves project
on all sides, square and pointed; the leaf-scars are rhombic, on
projecting leaf-cushions. The cones are pendulous. The cover-scales are
much shorter than the leathery, persisting ovuliferous scales.--The
genus _Larix_ (Larch) differs from all the others in having deciduous
leaves (the three preceding have leaves which persist for eleven to
twelve years). It has _long-branches_ with linear foliage-leaves and
short, thick, _perennial dwarf-branches_, which each year form a new
rosette of foliage-leaves, similar to those on the long-branches. The
male flowers and the erect cones resemble those of _Picea_, and are
borne on dwarf-branches.--_Cedrus_ (Cedar) resembles _Larix_ to some
extent, but has persistent leaves (_C. libani_, _C. deodara_).--_Pinus_
(Pine) has long-branches and dwarf-branches. The leaves of the
long-branches are scale-like and not green; the dwarf-branches have
very limited growth, and persist for three years; they arise in the
axils of the scales borne on the long-branches of the self-same year,
and each bears 2–5 foliage-leaves, they are also surrounded at the base
by a number of membranous bud-scales. The cone-scales have a _thick,
rhomboid extremity_ (the “shield”).
The buds which develope into long-branches arise at the apex of
other long-branches, and being very close together, form false
whorls. The female cones occupy the position of long-branches,
and take about two years for their development. The male flowers
arise close together, and form a spike-like inflorescence at
the base of a long-branch of the same year. The male flowers
occupy the position of dwarf-branches, so that a female cone
may be considered to be a modified long-branch, and a male cone
a modified dwarf-branch. The main axis of the seedling has
needle-like leaves, similar to those found on the older parts,
and on dwarf-branches; it is not until some time later that
the dwarf-branches are developed and the permanent arrangement
attained.
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