With regard to the Sympetalæ (or Gamopetalæ) it may be stated that
they form to a very great extent a closely connected and natural
group, having in common not only the character that the corolla is
gamopetalous and the stamens united with it (this being also found in
the Choripetalæ), but also a great many others (such as persistent
calyx, cyclic flowers with the formula S5, P5, A5 and as a rule G2,
the two carpels being united to form the ovary; seeds with a thick
integument and a very small nucellus). They are therefore considered
as an independent sub-class, and must be placed at the close of the
system of classification as the forms which presumably have arisen the
latest. In the future systems of classification this arrangement will
very probably be changed, and the first families of the Sympetalæ, the
Bicornes and others will for instance be to a certain extent united
with the families or orders of the Choripetalæ. The Sympetalæ may
certainly be considered as the youngest types, the strongly pronounced
metamorphosis supporting this theory, as also the formation of the
integument of the ovule, the one thick integument being undoubtedly
derived from the coalescence of two--a holochlamydeous ovule, etc.
The Apetalæ and Choripetalæ are united into one sub-class. The leaves
of the perianth in this case are, as a rule, free from each other, the
structure of the flowers presents many differences, and the ovules have
as a rule 2 integuments and a large nucellus. Considerable uncertainty
still prevails regarding the arrangement and the relationship of the
individual families of the Choripetalæ, and some of the following
families are hardly quite natural; but the best arrangement arrived at
so far has been adopted here.
At the beginning of the book a review of the orders of the Dicotyledons
will be found.
Sub-Class 1. =Choripetalæ. Petals free.=
Family 1. =Salicifloræ.=
Trees and shrubs, which, in the structure of the vegetative shoot and
the catkin-like inflorescences, resemble the Quercifloræ, but the
structure of the flower differs so much from them, that the only order
brought under this heading--_Salicaceæ_--well deserves to be separated
and to form a family of its own, the nearest relatives of which are
still doubtful. ~As Juglandaceæ and Myricaceæ also deserve to be placed
in a special family, the name _Amentaceæ_ (_Catkin-bearers_), hitherto
applied to all of these plants, cannot be retained as the name of a
family.~
[Illustration: FIG. 322.--Male and female catkins of _Salix
caprea_.]
There is only one order.
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