This family is on one side closely allied to the _Urticaceæ_ by its
solitary, _basal_, _vertical_, and _straight_ ovule, and by the conical
ocrea which envelopes the younger leaves in the bud, similar characters
being present in the Urticaceæ. On the other side it is related to
the Curvembryæ. The flowers are small, often _trimerous_, regular and
slightly perigynous (~in _Chloranthaceæ_, if they properly belong to
this family, and _Houttuynia_, more or less epigynous~). Syncarps are
present in some Piperaceæ, but the fruit is generally a single fruit,
one-seeded berry, nut or drupe. The leaves are generally scattered.
Order 1. =Polygonaceæ.= The majority are herbaceous plants with
round, often jointed stems, scattered leaves and _ocrea_, that is a
membranous, tubular, ligular or stipular structure _inside_ the base
of the leaf, which clasps the stem and axillary bud; the edges of the
lamina are rolled backwards in the bud. The flowers are regular, small,
generally ☿, slightly perigynous, with inconspicuous, simple, green or
white perianth of 5–6 free segments; stamens 5–9 (Fig. 354) sometimes
arranged in two series; gynœceum 2–3 carpels, ovary _unilocular_ with
_one basal_, _straight_ (orthotropous) _ovule_, 2–3 _free styles_.
The fruit is a 2–3-angular nut; the embryo, with mealy endosperm,
is straight or curved (Fig. 355 _H_), often unsymmetrical.--~The
inflorescences are compound, and generally branch from the axils of
the bracteoles, so that the last partial-inflorescences become coiled,
uniparous scorpioid cymes; in _Polygonum_ the two bracteoles unite into
a membranous tube; in _Rheum_ and _Rumex_ there is only one bracteole.~
[Illustration: FIG. 354.--_A_ Diagram of _Rheum_; _B_ of _Rumex_;
_C_ of _Polygonum fagopyrum_; _D_ of _P. lapathifolium_. The
ovules are indicated inside the ovaries; bracts and bracteoles
are not shown.]
_Rheum_ (Rhubarb, Fig. 354 _A_) has a 6-leaved, _petaloid_ perianth (Pn
3 + 3) and 9 stamens (A 3^2 + 3). The _3-winged_ nut is _not_ enclosed
by the perianth.
_Rumex_ (Dock, Fig. 354 _B_) has 6 stamens (A 3^2 + 0); the perianth is
6-leaved (Pr 3 + 3), green or red, and the triangular nut is enveloped
by the 3 interior perianth-leaves, which point upwards and continue to
grow after flowering. These perianth-leaves often have warts on their
outer surface. ~The following are monœcious: _R. acetosa_ and _R.
acetosella_.~
_Polygonum_ (Knot-grass, Figs. 354 _C_, _D_; 355). The _petaloid_
perianth is most frequently 5-merous (2/5 spiral); 5–8 stamens. The nut
is triangular (Fig. 354 _C_, 355), or lenticular (Fig. 354 _D_). ~There
are two whorls of stamens, the external with introrse, and the internal
with extrorse anthers. The gynœceum is often bicarpellate (Fig. 354
_D_).~
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