The flowers may be considered as constructed upon the
monocotyledonous type. _Pterostegia_ has a perfectly
monocotyledonous flower with 5 trimerous whorls. _Rheum_
likewise, but here the external staminal whorl is doubled (Fig.
254 _A_). _Oxyria_ has a dimerous _Rheum_-flower (4-leaved
perianth, 6 stamens, 2 stigmas). _Rumex_ has a _Rheum_-flower
with the suppression of the internal whorl of stamens (Fig.
354 _B_); _Emex_ is a dimerous _Rumex_. _Polygonum_, to which
_Coccoloba_, _Muehlenbeckia_ and others are related, differs
from _Rheum_ chiefly in having one of the leaves, which in the
latter takes part in the formation of the perianth, developed
in this case into a bracteole (so that the perianth is reduced
to five members), and several or all the stamens in the inner
whorl become suppressed.--The perianth in _Coccoloba_ and
_Muehlenbeckia_ is more or less perigynous and becomes fleshy,
enclosing the fruit. _Muehlenbeckia platyclada_ has flat branches
with rudimentary leaves; sometimes branches with normal,
arrow-shaped leaves are found. _Atraphaxis._
[Illustration: FIG. 355.--_Polygonum fagopyrum_: _A_ branch
with flower and fruits (nat. size); _B_ flower; _C_ the same
in longitudinal section; _D_ anterior and posterior view of
stamen; _E_ gynœceum; _F_ fruit (mag.); _G_ fruit in longitudinal
section; _H_ transverse section, showing the curved cotyledons
embedded in the endosperm; _I_ the embryo.]
POLLINATION. _Rumex_ is wind-pollinated, the stigmas are
therefore large and brush-like (indicated in Fig. 354 _B_).
_Rheum_ and _Polygonum_ are insect-pollinated and have therefore
capitate stigmas, etc.; honey-glands are situated at the base
of the stamens (_d_, in Fig. 354 _C_, and _n_ in Fig. 356); a
few small-flowered _Polygonum_ species are self-pollinated;
Buckwheat (_P. fagopyrum_) is dimorphic and has long-styled and
short-styled flowers (Fig. 356). _Pol. bistorta_ is protandrous
and homostyled.
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