The plants in this family have a _curved ovule_, and most frequently
a _kidney-shaped seed_ (generally provided with fine, cuticular,
projecting warts, Fig. 362 _B_), with a _curved, peripheral embryo
enclosing the endosperm which is most frequently floury_ (Figs. 362
_C_, 365 _H_; for exceptions, see Fig. 366); the seeds in all cases
are borne on a _centrally-placed_, and in most cases _free_, placenta
(they are “basal” when there is only 1 ovule in the ovary, Fig. 364).
The flower is regular, hypogynous or perigynous (Fig. 364) (only
rarely epigynous) and usually 5-_merous_. The flower which is most
complete has 5 whorls (S5, P5, A5+5, G2-3–5), as in some genera of
the Caryophyllaceæ (Figs. 360, 361); but from this type it becomes
reduced, the petals and stamens being suppressed, so that finally
5 perianth-leaves, 5 stamens (opposite the perianth-leaves), and 2
carpels (Fig. 361 _F_) only are present; for example, in certain genera
of the _Caryophyllaceæ_, in the _Chenopodiaceæ_, _Amarantaceæ_, and
others. When the number of stamens is increased to more than 5 in the
whorl, it is always possible to show that some of the stamens have been
divided. The number of the carpels and ovules also becomes reduced; in
the highest there is a central placenta, not free in its early stages,
with a large number of ovules; in those which are most reduced there is
only a single ovule, which is placed centrally at the base of the ovary
[Fig. 364]. Somewhat corresponding changes are found in the fruit,
which is a many-seeded _capsule_ in those which have many ovules, but
a one-seeded _nut_ where there is one ovule. In the most reduced forms
the flowers are generally unisexual.--Similar features are also present
in the vegetative parts. Almost all the species are herbaceous, the
leaves are simple and most frequently without stipules. ~The structure
of the stem, especially in Chenopodiaceæ, Amarantaceæ, Nyctaginiaceæ
and others, often differs from that of the ordinary Dicotyledon. In the
woody portion of the stem and root several rings are sometimes formed
which resemble annual rings but which are formed by new cambium-rings
arising outside the old ones which then cease to divide.~
Public-domain text, read in full here on John Shaqi.
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