Order 1. =Caryophyllaceæ.= Herbaceous plants, with round, nodose
stem; leaves _opposite_, slightly amplexicaul, simple, with sessile,
undivided, entire lamina; stipules nearly always absent; the
inflorescences are _dichasia_ passing over into unipared scorpioid
cymes. The flowers are regular, ☿ or unisexual, hypogynous or
perigynous, 5-(or 4-) merous with 2–3–4–5 carpels; calyx persistent;
corolla polypetalous. The ovary is unilocular (or originally, and
sometimes also in the later stages, plurilocular below, _e.g._
_Viscaria_), with _free styles_ and 1–several curved ovules on a
_central_, free placenta. The fruit is a nut or a capsule opening
apically with long or short valves (teeth, Fig. 362), equal to or
double the carpels. For the seeds refer to the family. ~In _Dianthus_
the embryo is straight.~
The flowers which are most complete have _Sn_, _Pn_, _An_ + _n_
(obdiplostemonous), _Gn_ where _n_ = 5 (Figs. 360, 361 _A_) or
4 (Fig. 361 _B_); the carpels may be placed opposite to the
sepals (Fig. 360) or opposite to the petals (Fig. 361 _A_, _B_).
Without any change taking place in the position of the other
whorls, the carpels are next found reduced to 2–3–4 (see the
genera); their number may easily be recognised by that of the
styles. This is the construction in the majority of the genera
in the two first groups. _Stellaria media_ differs considerably.
It may have (_a_) the flower as described above, with _G3_;
(_b_), the corolla only absent, or (_c_) only the petal-stamens
(A5 + 0, Fig. 361 _C_), or (_d_) all these as well as some of
the sepal-stamens. The same applies to _Sagina_, _Alsine_,
_Cerastium_, and others, and, finally, a series of genera are
formed, with certain conditions of reduction which have become
constant, and by a gradual series of steps lead to the most
reduced form, which has only 5 sepals and 5 (or even as far as
only 1) sepal-stamens (Fig. 361 _D_, _E_, _F_).--The PETALS in
the _Alsineæ_ are often deeply bifid. The sepal-stamens are most
frequently the longest, and bear nectaries at the base (Fig. 363
_st_). In the most complete forms the ovary has partition-walls
in the lower portion (Fig. 360); these do not, however, reach
to the top, and generally soon disappear. The ovules, when
numerous, are situated on the placenta in as many double rows as
there are carpels. In the number of ovules a reduction from many
to 1 takes place (Fig. 361). A comparison proves that the “free,
centrally placed” placenta is formed by the ventral portion of
the carpels. The single basal ovule in _Herniaria_ (Fig. 364),
_Scleranthus_, and others, is also borne on the carpels.
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