POLLINATION. _Alsineæ_ has ordinary nectaries at the base of
the calyx-stamens (Fig. 336): they are frequently protandrous
but may often, in the absence of cross-pollination (in the
less conspicuous species) pollinate themselves. Their open
flowers are accessible to many kinds of insects (particularly
flies and bees). _Gynodiœcious_ flowers are found in several
species, and the ☿-flowers are then generally more conspicuous
than the ♀-flowers. That the ♀-flowers have descended from
☿-flowers is seen by the large staminodes found in them (Fig.
363). _Arenaria peploides_ is diœcious (Fig. 363). The _Sileneæ_
are as a rule adapted for pollination by insects with long
probosces--especially butterflies,--and they are frequently
protandrous, so that at first the calyx-stamens open, later
on the corolla-stamens, then the stigmas expand. The honey is
secreted by a ring-like nectary round the base of the ovary or
by nectaries at the base of the stamens. Some only blossom and
emit scent at night or in the evening (_Lychnis vespertina_,
_Silene nutans_, _Saponaria officinalis_) and, like other
night-flowers, are of a white or pale colour.
DISTRIBUTION. 1,100 species, especially in temperate climates,
fewer in the colder zone, less still in the Tropics. The
Paronychieæ are especially found in dry, sandy fields.
USES. “Soap-root” (with _Saponin_, forming a lather in water)
from _Saponaria officinalis_ was formerly officinal, and
_Gypsophila struthium_. The seeds of _Agrostemma githago_ are
said to be poisonous.--The following are ornamental plants:
species of Pinks (_D. caryophyllus_, garden Pink, often
with double flowers; _D. barbatus_, _plumarius_, _etc._).
_Lychnis_, _Gypsophila_, _Silene_, _Cerastium_ (_C. tomentosum_
as edging for borders), _Saponaria officinalis_ (often
coronate).--_Spergula arvensis_ is sometimes cultivated.
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