[Illustration: FIG. 371.--Diagram of a dichasium of _Ranunculus
acer_: α_{1}, α^1, and β_{1}, β^1, bracteoles (the buds
in the axils of the bracteoles, α and α^1, are continued
antidromously). The flower has cyclic calyx and corolla, bub
acyclic (8/21) stamens.]
[Illustration: FIG. 372.--Diagram of an acyclic Ranunculaceous
flower (only 3 stamens are indicated). The spiral of the sepals
has a divergence of 3/5; that of the corolla and subsequent
leaves 3/8.]
Order 1. =Ranunculaceæ.= Nearly all are _herbs_ (except _Clematis_).
The leaves are scattered (except _Clematideæ_), they have a large
sheath with broad base (no stipules), and are most frequently
palminerved with palmate lobes. The flowers are hypogynous, with most
frequently a well pronounced convex receptacle (Figs. 374 _B_, 380), ☿,
regular (except _Delphinium_ and _Aconitum_); their structure varies
very much; in some the leaves are verticillate, in others arranged
spirally; in others, again, both modes of arrangement are found. It is
a characteristic feature that the various series of leaves (especially
calyx and corolla) are not so distinct or so sharply divided as
is usual. The leaves of the perianth are free, imbricate (except
_Clematideæ_); stamens _numerous_, with most frequently extrorse
anthers; gynœceum _free_, _apocarpous_ (except _Nigella_ and partly
_Helleborus_), with 1 or several ovules (Figs. 373, 378, 379) borne on
the ventral suture. The fruit is either a nut or a follicle (_Actæa_
has berries). The seed has a _large, oil-containing_ endosperm and a
small embryo (Fig. 374).
The main axis generally terminates in a flower, and the
lateral axes branch in a cymose manner (Fig. 371). The flowers
show the following differences in construction: VERTICILLATE
(EUCYCLIC), _i.e._ constructed all through of alternating
whorls: _Aquilegia_ (Fig. 370), _Xanthorhiza_, and sometimes
_Eranthis_. SEMIVERTICILLATE (HEMICYCLIC) _i.e._ with sepals and
petals in alternate whorls, and the others arranged spirally:
_Ranunculus_ (Fig. 371), _Myosurus_, _Pæonia_ and several other
genera entirely, or in certain species only. SPIRAL-FLOWERED
(ACYCLIC) _i.e._ all the leaves are arranged spirally, so
that sepals and petals do not alternate the one with the
other, even though they are the same in number: _Adonis_ (Fig.
372), _Aconitum_, _Delphinium_-species, _Nigella_-species,
_Helleborus_. The leaves of the calyx are in this instance
arranged on a spiral of 2/5; those of the corolla on 2/5,
3/8, 5/13 or 8/21, and stamens and carpels likewise on higher
fractions of the same series.
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