The genera _Caltha_, _Anemone_, _Thalictrum_ and _Clematis_
have a _single perianth_, which is most frequently petaloid;
it is thus apparent that the sepals are petaloid, and the
leaves, which in other genera have developed as petals, are in
these instances stamens. The calyx is similarly petaloid in
the genera _Helleborus_, _Eranthis_, _Nigella_, _Delphinium_
and _Aconitum_; but the petals are present in these instances
in unusual (horn-like) forms, and almost entirely given
up to the function of nectaries, a function they already
possess in _Ranunculus_. According to a more recent theory
the “honey-leaves” are transformed stamens, which have lost
the function of reproduction; the perianth is then single,
and most frequently petaloid. [Those leaves in the flowers of
many Ranunculaceæ which bear nectaries are termed by Prantl
honey-leaves, and comprise those leaf-structures of the flower
whose essential function lies in the production of nectar,
and which, independent of the differentiation of the perianth
into calyx and corolla, are derived from the stamens by the
loss of their reproductive functions. Clear transitional
forms are found between the two series of the perianth
(_e.g._ between the sepaloid and petaloid perianth-leaves
of _Anemone japonica_, _A. decapetala_, _Trollius_-species)
while transitional forms are never found between perianth-and
honey-leaves (with the exception of _Aquilegia vulgaris_, var.
_stellata_). In _Anemone_ and _Clematis_ the honey-leaves
pass gradually into the stamens, and agree with the stamens
in the other Ranunculaceæ in their arrangement, development,
and scant system of veins (except _Nigella_). In _Delphinium_,
sect. _Consolida_, the two honey-leaves placed in front of
the unpaired perianth-leaf are united into one, as shown by
the veins (twice three veins arranged symmetrically). The
honey-leaves of _Aquilegia_, _Callianthemum_, and the majority
of the _Ranunculus_-species serve by reason of their large
circumference, as organs of attraction, and on this account
are considered as petals by other authors.--The same position
in the flower which the honey-leaves assume is found occupied
by staminodes, without nectar, in some _Coptis_-species, in
_Anemonopsis_, _Actæa_ sect. _Euactæa_, (_e.g. A. racemosa_),
_Clematis_ sect. _Atragene_; in the last-named they closely
surround the stamens, in _Actæa_ they are petaloid.--A perianth,
sharply differentiated into calyx and corolla, and destitute of
honey-leaves, is found in _Anemone_, sect. _Knowltonia_ (Cape),
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