_Adonis_, _Pæonia_.--The perianth of the Ranunculaceæ is
considered by Prantl to be usually petaloid.--The nectaries
arise in the Ranunculaceæ (1) on normal stamens (_Clematis_
sect. _Viorna_), (2) on the honey-leaves (this is generally the
case), and (3) on the carpels (_Caltha_ and the majority of
_Trollius_-species).--As the result of his researches upon the
Ranunculaceæ, Prantl does not agree with the view advanced by
Drude (Schenk, _Hand. d. Bot._ iii.) that the petals in general
have proceeded from the metamorphosis of the stamens (_K_)].
[Illustration: FIG. 373.--Ovaries in longitudinal section: _v_
the ventral suture; _d_ the dorsal suture: _A_, _B Clematis_;
_C Ranunculus_; _D Myosurus_.]
The most primitive form of fruit is undoubtedly the pod formed
by one carpel, on the edges of which (along the ventral suture)
two rows of ovules are situated: Pæonieæ, Helleboreæ, Delphinieæ
(Fig. 379). In a great many genera the number of ovules has been
limited to _one_ perfect one, which is placed in the central
plane under the united leaf-edges, and sometimes also some
barren ovules above it (Fig. 373). The fruitlets in this case
become achenes, and are present in much larger numbers than when
there are follicles.
[Illustration: FIG. 374.--_Helleborus niger_: _A_ flower; _B_
receptacle; _pet_ petals (honey-leaves); _pi_ stamens and
carpels; _C_ seed; _D_ anther (cross section); _alb_ endosperm.]
[Illustration: FIG. 375.--_Caltha palustris_: fruit.]
The following have FOLLICLES: _Pæonieæ_, _Helleboreæ_ (except _Actæa_)
and _Delphinieæ_; ACHENES: _Ranunculeæ_, _Anemoneæ_ and _Clematideæ_.
=A. Follicles= (Figs. 375, 379), with many ovules, situated in two rows
along the ventral suture. ~_Actæa_ has berries, _Nigella_ has capsules
of several loculi.~
=1.= PÆONIEÆ, PEONY GROUP. This has regular, acyclic flowers with a
normal, most frequently 5-leaved, imbricate calyx; large, coloured
petals, and introrse anthers. Slightly perigynous. Surrounding the
base of the carpels a ring-like swelling of the receptacle (“disc”) is
present, which is largest in _P. moutan_. The follicles are more or
less fleshy or leathery. Mostly herbs, with pinnatisect or decompound
leaves and large, solitary flowers; a gradual transition may be traced
from the foliage-leaves to the petals. _Pæonia; Hydrastis._
[Illustration: FIG. 376.--_Aquilegia vulgaris._]
[Illustration: FIG. 377.--_Caltha palustris_ (nat. size).]
[Illustration: FIG. 378.--_Nigella_: _A_, _B_ fruit of _N.
damascena_, entire, and cut transversely. _C_ Petal (honey-leaf)
of _N. arvensis_. _D_ Petal of _N. damascena_.]
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